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fissidens moss, yew-leaf pocket moss

pale-nerve fissidens moss

Habit Plants to 11 × 3.5 mm. Plants to 6 × 1 mm.
Stem

mostly branched;

axillary hyaline nodules absent;

central strand present.

branched and unbranched;

axillary hyaline nodules absent;

central strand weak or absent.

Leaves

as many as 15 pairs, often undulate, oblong to lanceolate, obtuse to broadly acute, cuspidate, to 2.7 mm × 0.6 mm;

dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent;

vaginant laminae 2/3 the leaf length, ± equal, minor lamina ending on or near margin;

margin evenly serrulate, often crenulate-serrulate on dorsal and ventral laminae, elimbate;

costa ending in cuspidate apex, taxifolius-type;

laminal cells 1-stratose, distinct, smooth, bulging, firm-walled, irregularly hexagonal, 8–11 µm frequently paler at margin, conspicuously mammillose in vaginant laminae, larger and clearer juxtacostally.

as many as 18 pairs, lingulate to lanceolate, rounded to obtuse to broadly acute, to 1 × 0.25 mm;

dorsal lamina narrowed or rounded proximally, ending at insertion, not decurrent;

vaginant laminae ± 1/2–2/3 leaf length, ± unequal, minor lamina ending near margin;

margin serrulate, limbate, ± entire in proximal 1/3–1/2 of perichaetial and subtending 1–2 pairs of leaves, limbidium sometimes indistinct, limbidial cells 1-stratose;

costa ending 3–25 cells before apex, often spurred distally, bryoides-type;

laminal cells 1-stratose, pluripapillose, obscure, firm-walled, irregularly quadrate to hexagonal, 4–8 µm, in transverse section usually twice as deep as wide.

Seta

to 17 mm.

to 2 mm.

Sexual condition

rhizautoicous;

perigonia on short branches proximal to elongate stems;

perichaetia on short, proximal, axillary branches.

rhizautoicous, cladautoicous, rarely gonioautoicous.

Capsule

theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.5 mm;

peristome taxifolius-type;

operculum to 1 mm.

theca exserted, erect, radially symmetric, to 0.6 mm;

peristome scariosus-type;

operculum 0.3 mm.

Calyptra

cucullate, smooth, to 2 mm.

cucullate, ± prorate, 0.3 µm.

Spores

13–18 µm.

9–14µm.

Sporophytes

1 per perichaetium.

1–2 per perichaetium.

Fissidens taxifolius

Fissidens pallidinervis

Habitat Damp, shaded soil, humus, and rocks Usually in damp areas around bases of trees, decaying logs, occasionally limestone and soil
Distribution
from FNA
AL; AR; AZ; CA; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; SC; TN; TX; VA; VT; WA; WI; WV; NS; NT; ON; PE; QC; Mexico; Central America; West Indies; South America (Brazil and Chile); Europe; Asia (China, Japan); Pacific Islands (New Zealand)
[WildflowerSearch map]
from FNA
FL; LA; Mexico; Central America; South America; West Indies; Asia; Africa
Discussion

Fissidens taxifolius is distinguished by evenly serrulate or crenulate-serrulate leaf margin, stout costa ending in leaf cusp, conspicuous mammillose cells in the vaginant laminae, and sporophytes on short branches in the axils of proximal leaves. It can be confused with F. bushii, but that species is smaller, has costa shorter, and cells in the vaginant laminae have small, more or less inconspicuous papillae in the corners of the walls. Fissidens taxifolius usually occurs on shaded, damp soil or humus, whereas F. bushii is usually found on disturbed soil in woods, along paths, and in road cuts. Fissidens clebschii is considered to be a freakish expression of F. taxifolius (R. A. Pursell 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens pallidinervis and F. elegans are the only species of the genus in North America with small, obscure, pluripapillose laminal cells. Fissidens pallidinervis, however, is distinguished by a rounded to broadly acute leaf apex, and a limbidium restricted to the lower parts of vaginant laminae of perichaetial and one or two pairs of subtending leaves. Axillary, stalked, multicellular, clavate gemmae have been reported in Japanese collections of F. pallidinervis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 341. FNA vol. 27, p. 350.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. clebschii F. garberi, F. minutus
Name authority Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 1–5. (1801) Mitten: J. Linn. Soc., Bot. 12: 592. (1869)
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