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fissidens moss, yew-leaf pocket moss

little's fissidens moss

Habit Plants to 11 × 3.5 mm. Plants to 3 × 2 mm.
Stem

mostly branched;

axillary hyaline nodules absent;

central strand present.

unbranched and sparingly branched;

axillary hyaline nodules absent;

central strand absent.

Leaves

as many as 15 pairs, often undulate, oblong to lanceolate, obtuse to broadly acute, cuspidate, to 2.7 mm × 0.6 mm;

dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent;

vaginant laminae 2/3 the leaf length, ± equal, minor lamina ending on or near margin;

margin evenly serrulate, often crenulate-serrulate on dorsal and ventral laminae, elimbate;

costa ending in cuspidate apex, taxifolius-type;

laminal cells 1-stratose, distinct, smooth, bulging, firm-walled, irregularly hexagonal, 8–11 µm frequently paler at margin, conspicuously mammillose in vaginant laminae, larger and clearer juxtacostally.

as many as 8 pairs, ligulate, obtuse-apiculate to acute, to 1.5 × 0.25 mm;

dorsal lamina narrowed proximally, ending at insertion or slightly decurrent;

vaginant laminae 1/2–2/3 leaf length, ± equal, minor lamina ending on or near margin;

margin crenulate, often irregularly so on vaginant laminae, elimbate;

costa ending 3–5 cells before apex, bryoides-type;

lamina cells 1-stratose, distinct, mammillose, firm-walled, irregularly quadrate to hexagonal, oblate on margin, 8–10 µm, somewhat larger, ± oblong in proximal parts of vaginant laminae.

Seta

to 17 mm.

1.4–1.8 mm.

Sexual condition

rhizautoicous;

perigonia on short branches proximal to elongate stems;

perichaetia on short, proximal, axillary branches.

rhizautoicous and gonio-autoicous;

perigonia gemmiform and on elongate stems.

Capsule

theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.5 mm;

peristome taxifolius-type;

operculum to 1 mm.

theca exserted, erect, radially symmetric, to 0.5 mm long;

peristome teeth anomalous, undivided, papillose;

operculum to 0.3 mm.

Calyptra

cucullate, smooth, to 2 mm.

cucullate, smooth, 0.3 mm.

Spores

13–18 µm.

8–11 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens taxifolius

Fissidens littlei

Habitat Damp, shaded soil, humus, and rocks Walls of small pockets in gypsum sinkholes
Distribution
from FNA
AL; AR; AZ; CA; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; SC; TN; TX; VA; VT; WA; WI; WV; NS; NT; ON; PE; QC; Mexico; Central America; West Indies; South America (Brazil and Chile); Europe; Asia (China, Japan); Pacific Islands (New Zealand)
[WildflowerSearch map]
from FNA
NM
Discussion

Fissidens taxifolius is distinguished by evenly serrulate or crenulate-serrulate leaf margin, stout costa ending in leaf cusp, conspicuous mammillose cells in the vaginant laminae, and sporophytes on short branches in the axils of proximal leaves. It can be confused with F. bushii, but that species is smaller, has costa shorter, and cells in the vaginant laminae have small, more or less inconspicuous papillae in the corners of the walls. Fissidens taxifolius usually occurs on shaded, damp soil or humus, whereas F. bushii is usually found on disturbed soil in woods, along paths, and in road cuts. Fissidens clebschii is considered to be a freakish expression of F. taxifolius (R. A. Pursell 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

This rare Fissidens, named in honor of the American botanist Elbert Little, is known only from the type locality and was recently re-discovered there by K. W. Allred (1998). The plants are pale green, probably the result of their occurrence in small pocketlike depressions where direct sunlight does not penetrate. The species has been confused with F. amoenus, but differs from the latter by its elimbate leaves, slightly smaller, mammillose laminal cells, oblate marginal cells, absence of enlarged pellucid cells in the proximal parts of the vaginant laminae, and undivided, papillose peristome teeth.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 341. FNA vol. 27, p. 354.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. clebschii Moenkemeyera littlei
Name authority Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 1–5. (1801) (R. S. Williams) Grout: Moss Fl. N. Amer. 1: 249. (1939)
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