Fissidens taxifolius |
Fissidens bushii |
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fissidens moss, yew-leaf pocket moss |
Bush's fissidens moss |
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Habit | Plants to 11 × 3.5 mm. | Plants to 13 × 2–2.5 mm. |
Stem | mostly branched; axillary hyaline nodules absent; central strand present. |
branched; axillary hyaline nodules weak; central strand weak. |
Leaves | as many as 15 pairs, often undulate, oblong to lanceolate, obtuse to broadly acute, cuspidate, to 2.7 mm × 0.6 mm; dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent; vaginant laminae 2/3 the leaf length, ± equal, minor lamina ending on or near margin; margin evenly serrulate, often crenulate-serrulate on dorsal and ventral laminae, elimbate; costa ending in cuspidate apex, taxifolius-type; laminal cells 1-stratose, distinct, smooth, bulging, firm-walled, irregularly hexagonal, 8–11 µm frequently paler at margin, conspicuously mammillose in vaginant laminae, larger and clearer juxtacostally. |
as many as 20 pairs, oblong to lanceolate, obtuse-apiculate to acute-apiculate, to 2 × 0.5 mm; dorsal lamina rounded proximally, ending at insertion, not decurrent; vaginant laminae ± 2/3 leaf length, ± equal, minor lamina ending on or near margin; margin crenulate-serrulate, more strongly serrulate on vaginant laminae; costa ending 1–2 cells before apex or ending in apiculus, taxifolius-type; laminal cells irregularly 2-stratose in dorsal and ventral laminae, smooth, bulging, firm-walled, rounded-hexagonal, 7–9 µm, papillose in cell corners with 2–4 ± inconspicuous papillae in vaginant laminae. |
Seta | to 17 mm. |
to 10 mm. |
Sexual condition | rhizautoicous; perigonia on short branches proximal to elongate stems; perichaetia on short, proximal, axillary branches. |
dioicous; perigonia and perichaetia gemmiform, axillary, confined to proximal parts of stems. |
Capsule | theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum to 1 mm. |
theca to 1 mm, erect, ± arcuate, bilaterally symmetric, about 1 mm; peristome taxifolius-type; operculum 1 mm. |
Calyptra | cucullate, smooth, to 2 mm. |
cucullate, smooth, 0.6 mm. |
Spores | 13–18 µm. |
14–18 µm. |
Sporophytes | 1 per perichaetium. |
1 per perichaetium. |
Fissidens taxifolius |
Fissidens bushii |
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Habitat | Damp, shaded soil, humus, and rocks | Usually on bare or disturbed clayey soil in open and in woods, along paths, roadside banks, along streams, ravines, infrequently on rocks, stones, tree bases |
Distribution |
AL; AR; AZ; CA; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; SC; TN; TX; VA; VT; WA; WI; WV; NS; NT; ON; PE; QC; Mexico; Central America; West Indies; South America (Brazil and Chile); Europe; Asia (China, Japan); Pacific Islands (New Zealand)
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AL; AR; DC; DE; FL; GA; IA; IL; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; VT; WV; NB; NS; ON; QC |
Discussion | Fissidens taxifolius is distinguished by evenly serrulate or crenulate-serrulate leaf margin, stout costa ending in leaf cusp, conspicuous mammillose cells in the vaginant laminae, and sporophytes on short branches in the axils of proximal leaves. It can be confused with F. bushii, but that species is smaller, has costa shorter, and cells in the vaginant laminae have small, more or less inconspicuous papillae in the corners of the walls. Fissidens taxifolius usually occurs on shaded, damp soil or humus, whereas F. bushii is usually found on disturbed soil in woods, along paths, and in road cuts. Fissidens clebschii is considered to be a freakish expression of F. taxifolius (R. A. Pursell 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens bushii, named in honor of the collector of the type, Benjamin Franklin Bush, is most apt to be confused with F. taxifolius. However, plants of F. bushii are usually much smaller, costa shorter, and leaves sometimes caducous. Moreover, small papillae restricted to the corners of cells in vaginant laminae are distinctive. The species is probably most closely related to the Asiatic F. teysmannianus Dozy & Molkenboer, which also has inconspicuous papillae in the corners of vaginant laminal cells. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 341. | FNA vol. 27, p. 342. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. clebschii | F. subbasilaris var. bushii |
Name authority | Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 1–5. (1801) | (Cardot & Thériot) Cardot & Thériot: Bot. Gaz. 37: 365. (1904) |
Web links |