Fissidens taxifolius |
Fissidens arcticus |
|
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fissidens moss, yew-leaf pocket moss |
arctic fissidens moss |
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Habit | Plants to 11 × 3.5 mm. | Plants to 14 × 3 mm. |
Stem | mostly branched; axillary hyaline nodules absent; central strand present. |
usually unbranched; axillary hyaline nodules absent; central strand weak. |
Leaves | as many as 15 pairs, often undulate, oblong to lanceolate, obtuse to broadly acute, cuspidate, to 2.7 mm × 0.6 mm; dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent; vaginant laminae 2/3 the leaf length, ± equal, minor lamina ending on or near margin; margin evenly serrulate, often crenulate-serrulate on dorsal and ventral laminae, elimbate; costa ending in cuspidate apex, taxifolius-type; laminal cells 1-stratose, distinct, smooth, bulging, firm-walled, irregularly hexagonal, 8–11 µm frequently paler at margin, conspicuously mammillose in vaginant laminae, larger and clearer juxtacostally. |
as many as 40 pairs, lanceolate to oblong-lanceolate, generally broadly acute, 0.4–1.5 × 0.2–0.3 mm; dorsal laminae narrowed proximally, often ending well before insertion; vaginant laminae 1/2–3/4 leaf length, ± equal, minor lamina ending on or near margin; margin entire but often ± serrate distally, limbate on the vaginant laminae, infrequently limbate on dorsal and ventral laminae of perichaetial leaves, limbidial cells 1-stratose; costa usually ending 2–3 cells before apex, bryoides-type; laminal cells 1-stratose, distinct, smooth, slightly bulging, firm-walled, irregularly quadrate to hexagonal, 8–11 µm. |
Seta | to 17 mm. |
1.75–6 mm. |
Sexual condition | rhizautoicous; perigonia on short branches proximal to elongate stems; perichaetia on short, proximal, axillary branches. |
polyoicous, most frequently gonioautoicous; naked antheridia often in axils of distal leaves. |
Capsule | theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum to 1 mm. |
theca exserted, usually erect, radially symmetric, 0.4–0.8 mm; peristome bryoides-type; operculum 0.6 mm. |
Calyptra | cucullate, smooth, to 2 mm. |
not seen. |
Spores | 13–18 µm. |
14–20 µm. |
Sporophytes | 1 per perichaetium. |
1 per perichaetium. |
Fissidens taxifolius |
Fissidens arcticus |
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Habitat | Damp, shaded soil, humus, and rocks | Bare soil in small tufts in wet meadows, silt in frost boils, rock crevices, also as slender strands among tuft-forming bryophytes |
Distribution |
AL; AR; AZ; CA; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; SC; TN; TX; VA; VT; WA; WI; WV; NS; NT; ON; PE; QC; Mexico; Central America; West Indies; South America (Brazil and Chile); Europe; Asia (China, Japan); Pacific Islands (New Zealand)
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AK; NT; NU; YT; Greenland; n Europe |
Discussion | Fissidens taxifolius is distinguished by evenly serrulate or crenulate-serrulate leaf margin, stout costa ending in leaf cusp, conspicuous mammillose cells in the vaginant laminae, and sporophytes on short branches in the axils of proximal leaves. It can be confused with F. bushii, but that species is smaller, has costa shorter, and cells in the vaginant laminae have small, more or less inconspicuous papillae in the corners of the walls. Fissidens taxifolius usually occurs on shaded, damp soil or humus, whereas F. bushii is usually found on disturbed soil in woods, along paths, and in road cuts. Fissidens clebschii is considered to be a freakish expression of F. taxifolius (R. A. Pursell 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Differing opinions have been expressed on the taxonomic standing of Fissidens arcticus. According to W. C. Steere (1978), it might be only an environmentally induced expression of F. bryoides, but A. A. Frisvoll (1981), who reported the species from Svalbard, suggested that it be treated as a subspecies or variety of F. bryoides. They and other previous workers (Steere and G. R. Brassard 1974; M. A. Bruggeman-Nannenga and E. Nyholm 1986), however, did not mention the dimorphic stems, of which the fertile (perichaetial) ones are shorter and have fewer pairs of leaves than the infertile ones. That character and the tendency of the limbidium to be expressed on all laminae of the leaves of perichaetial stems but otherwise restricted to the vaginant lamina of infertile stems distinguish F. arcticus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 341. | FNA vol. 27, p. 346. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. clebschii | |
Name authority | Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 1–5. (1801) | Bryhn: Bryoph. Itin. Pol. Norv., 57, plate 1, fig. 2. (1906) |
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