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fissidens moss

Habit Plants to 12 × 1.5–3.5 mm. Plants to 5 × 2 mm.
Stem

branched;

axillary hyaline nodules absent;

central strand present.

unbranched and branched;

axillary hyaline nodules absent;

central strand weak.

Leaves

as many as 28 pairs, sometimes ± undulate, oblong, obtuse-apiculate to acute, to 2.4 × 0.3–0.4 mm;

dorsal lamina narrowed proximally, ending before or at insertion;

vaginant laminae 1/2–2/3 leaf length, acute, ± unequal, minor lamina ending near margin;

margin evenly crenulate-serrulate but irregularly serrate distally;

costa ending 6–16 cells before apex, covered and obscured above by chlorophyllose cells, taxifolius-type;

lamina cells irregularly 2-stratose, smooth, bulging, firm-walled, irregularly rounded-hexagonal, 7.5–10 µm. Sexual condition gonioautoicous and cladautoicous;

perigonia gemmiform, axillary, and on short branches;

perichaetia on short axillary branches near proximal ends of stem.

as many as 10 pairs, lanceolate to lingulate, acute to obtuse-apiculate, to 1.3 × 0.4 mm;

dorsal lamina narrowed abruptly proximally, ending at insertion, not decurrent;

vaginant laminae ± 1/2 leaf length, equal;

margin serrulate, limbate and entire on complete length of vaginant laminae and often on proximal portion of adjacent ventral laminae of all leaves except smallest, limbidial cells 1-stratose;

costa percurrent to ending 2–3 cells before apex, bryoides-type;

laminal cells 1-stratose, distinct, mammillose, firm-walled, quadrate to hexagonal, 7–9 µm. Sexual condition rhizautoicous;

perigonia gemmiform, proximal to fertile and infertile stems.

Seta

1.5–5.5 mm.

to 3.5 mm.

Capsule

theca 0.8–2.1 mm, erect, radially symmetric or nearly so;

peristome taxifolius-type;

operculum long rostrate, 0.6–0.9 mm.

theca exserted, erect, radially symmetric, to 0.6 mm;

peristome scariosus-type;

operculum to 9.4 mm.

Calyptra

cucullate, smooth, 1.5 mm.

cucullate, smooth, to 0.5 mm.

Spores

13–18 µm.

11–23 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens subbasilaris

Fissidens submarginatus

Habitat Particularly around bases of trees, also rocks, mostly limestone, infrequently decaying wood Moist, disturbed soil and around bases of trees, mostly in cypress swamps
Distribution
from FNA
AL; AR; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; QC; Mexico (San Luis Potosí, Tamaulipas)
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from FNA
LA; Mexico; Central America; South America; West Indies; Africa
Discussion

Fissidens subbasilaris is distinguished, even in the field, by a costa obscured by chlorophyllose cells distally, and short perichaetial branches in the axils of proximal leaves. The plants are typically dark-green and often tinged with red.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens submarginatus is most likely to be confused with F. leptophyllus. Both species have mammillose laminal cells, serrulate leaf margin, and limbidium of 1-stratose cells restricted essentially to vaginant laminae. The limbidium is the best character by which these two species are separated. In F. submarginatus the limbidium consistently extends the entire length of the vaginant laminae of most or all leaves, and on occasion extends a short distance onto the proximal part of the adjacent ventral lamina. In contrast, the limbidium in F. leptophyllus consistently extends no more than 1/2 the length of the vaginant laminae of most leaves. Minor differences can also be found in the shape of the leaf apex (usually apiculate vs. not apiculate), costa length (ending 2–3 cells before the leaf apex vs. ending 2–6 cells before the leaf apex), and cell size (7–9 µm versus 7–12 µm).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 342. FNA vol. 27, p. 352.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. kochii
Name authority Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 6–9. (1801) Bruch: Flora 29: 133. (1846)
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