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fissidens moss

fissidens moss

Habit Plants to 12 × 1.5–3.5 mm. Plants to 3.5 × 3 mm.
Stem

branched;

axillary hyaline nodules absent;

central strand present.

unbranched;

axillary hyaline nodules absent;

central strand absent.

Leaves

as many as 28 pairs, sometimes ± undulate, oblong, obtuse-apiculate to acute, to 2.4 × 0.3–0.4 mm;

dorsal lamina narrowed proximally, ending before or at insertion;

vaginant laminae 1/2–2/3 leaf length, acute, ± unequal, minor lamina ending near margin;

margin evenly crenulate-serrulate but irregularly serrate distally;

costa ending 6–16 cells before apex, covered and obscured above by chlorophyllose cells, taxifolius-type;

lamina cells irregularly 2-stratose, smooth, bulging, firm-walled, irregularly rounded-hexagonal, 7.5–10 µm. Sexual condition gonioautoicous and cladautoicous;

perigonia gemmiform, axillary, and on short branches;

perichaetia on short axillary branches near proximal ends of stem.

as many as 6 pairs, lanceolate, acute to obtuse, sometimes apiculate, to 2.2 × 0.5 mm;

dorsal lamina narrowed proximally, ending at insertion, not decurrent;

vaginant laminae 1/3–1/2 leaf length, equal;

margin entire but sometimes ± serrulate distally, limbate, limbidium 1–2 cells wide, ending before apex, often indistinct on vaginant laminae, limbidial cells 1-stratose; ecostate, or costa consisting of very short proximal vestige;

laminal cells 1-stratose, smooth, thin-walled, hexagonal to elongate-hexagonal, 31–67 × 14–34 µm, ± oblong in proximal portions of leaves, particularly in vaginant laminae.

Seta

1.5–5.5 mm.

to 3 mm.

Sexual condition

rhizautoicous;

perigonia gemmiform, proximal to perichaetial stems, and on elongate stems, to 1.2 mm.

Capsule

theca 0.8–2.1 mm, erect, radially symmetric or nearly so;

peristome taxifolius-type;

operculum long rostrate, 0.6–0.9 mm.

theca exserted, erect, radially symmetric, sometimes slightly arcuate, bilaterally symmetric, to 0.4 mm;

peristome scariosus-type;

operculum ± as long as theca.

Calyptra

cucullate, smooth, 1.5 mm.

mitrate, prorate, 0.4 mm.

Spores

13–18 µm.

9–13 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens subbasilaris

Fissidens hyalinus

Habitat Particularly around bases of trees, also rocks, mostly limestone, infrequently decaying wood Bare, clayey soil in shaded ravines, gorges, and dry stream bottoms
Distribution
from FNA
AL; AR; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; QC; Mexico (San Luis Potosí, Tamaulipas)
[WildflowerSearch map]
from FNA
KY; LA; OH; PA; TN; Asia
Discussion

Fissidens subbasilaris is distinguished, even in the field, by a costa obscured by chlorophyllose cells distally, and short perichaetial branches in the axils of proximal leaves. The plants are typically dark-green and often tinged with red.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens hyalinus (Latin hyalus, literally “of glass,” alluding to the transparent, glasslike appearance) is doubtless more common than previously thought. B. E. Lemmon (1966) published a photograph showing a perigonial stem and a perichaetial stem with attached sporophyte. A. C. Risk (2002) gave a thorough account of the distribution and habitat of this tiny moss. Plants cannot be mistaken for any other species in North America, owing to the ecostate leaves and smooth, large, thin-walled laminal cells that shrink considerably when dry.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 342. FNA vol. 27, p. 357.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Name authority Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 6–9. (1801) Wilson & Hooker f.: J. Bot. (Hooker) 3: 89, fig. 2. (1841)
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