Fissidens subbasilaris |
Fissidens arcticus |
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fissidens moss |
arctic fissidens moss |
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Habit | Plants to 12 × 1.5–3.5 mm. | Plants to 14 × 3 mm. |
Stem | branched; axillary hyaline nodules absent; central strand present. |
usually unbranched; axillary hyaline nodules absent; central strand weak. |
Leaves | as many as 28 pairs, sometimes ± undulate, oblong, obtuse-apiculate to acute, to 2.4 × 0.3–0.4 mm; dorsal lamina narrowed proximally, ending before or at insertion; vaginant laminae 1/2–2/3 leaf length, acute, ± unequal, minor lamina ending near margin; margin evenly crenulate-serrulate but irregularly serrate distally; costa ending 6–16 cells before apex, covered and obscured above by chlorophyllose cells, taxifolius-type; lamina cells irregularly 2-stratose, smooth, bulging, firm-walled, irregularly rounded-hexagonal, 7.5–10 µm. Sexual condition gonioautoicous and cladautoicous; perigonia gemmiform, axillary, and on short branches; perichaetia on short axillary branches near proximal ends of stem. |
as many as 40 pairs, lanceolate to oblong-lanceolate, generally broadly acute, 0.4–1.5 × 0.2–0.3 mm; dorsal laminae narrowed proximally, often ending well before insertion; vaginant laminae 1/2–3/4 leaf length, ± equal, minor lamina ending on or near margin; margin entire but often ± serrate distally, limbate on the vaginant laminae, infrequently limbate on dorsal and ventral laminae of perichaetial leaves, limbidial cells 1-stratose; costa usually ending 2–3 cells before apex, bryoides-type; laminal cells 1-stratose, distinct, smooth, slightly bulging, firm-walled, irregularly quadrate to hexagonal, 8–11 µm. Sexual condition polyoicous, most frequently gonioautoicous; naked antheridia often in axils of distal leaves. |
Seta | 1.5–5.5 mm. |
1.75–6 mm. |
Capsule | theca 0.8–2.1 mm, erect, radially symmetric or nearly so; peristome taxifolius-type; operculum long rostrate, 0.6–0.9 mm. |
theca exserted, usually erect, radially symmetric, 0.4–0.8 mm; peristome bryoides-type; operculum 0.6 mm. |
Calyptra | cucullate, smooth, 1.5 mm. |
not seen. |
Spores | 13–18 µm. |
14–20 µm. |
Sporophytes | 1 per perichaetium. |
1 per perichaetium. |
Fissidens subbasilaris |
Fissidens arcticus |
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Habitat | Particularly around bases of trees, also rocks, mostly limestone, infrequently decaying wood | Bare soil in small tufts in wet meadows, silt in frost boils, rock crevices, also as slender strands among tuft-forming bryophytes |
Distribution |
AL; AR; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; QC; Mexico (San Luis Potosí, Tamaulipas)
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AK; NT; NU; YT; Greenland; n Europe |
Discussion | Fissidens subbasilaris is distinguished, even in the field, by a costa obscured by chlorophyllose cells distally, and short perichaetial branches in the axils of proximal leaves. The plants are typically dark-green and often tinged with red. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Differing opinions have been expressed on the taxonomic standing of Fissidens arcticus. According to W. C. Steere (1978), it might be only an environmentally induced expression of F. bryoides, but A. A. Frisvoll (1981), who reported the species from Svalbard, suggested that it be treated as a subspecies or variety of F. bryoides. They and other previous workers (Steere and G. R. Brassard 1974; M. A. Bruggeman-Nannenga and E. Nyholm 1986), however, did not mention the dimorphic stems, of which the fertile (perichaetial) ones are shorter and have fewer pairs of leaves than the infertile ones. That character and the tendency of the limbidium to be expressed on all laminae of the leaves of perichaetial stems but otherwise restricted to the vaginant lamina of infertile stems distinguish F. arcticus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 342. | FNA vol. 27, p. 346. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Name authority | Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 6–9. (1801) | Bryhn: Bryoph. Itin. Pol. Norv., 57, plate 1, fig. 2. (1906) |
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