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fissidens moss

Appalachian fissidens moss

Habit Plants to 12 × 1.5–3.5 mm. Plants to 13 × 2–4 mm.
Stem

branched;

axillary hyaline nodules absent;

central strand present.

unbranched and branched;

axillary hyaline nodules weak or absent;

central strand weak.

Leaves

as many as 28 pairs, sometimes ± undulate, oblong, obtuse-apiculate to acute, to 2.4 × 0.3–0.4 mm;

dorsal lamina narrowed proximally, ending before or at insertion;

vaginant laminae 1/2–2/3 leaf length, acute, ± unequal, minor lamina ending near margin;

margin evenly crenulate-serrulate but irregularly serrate distally;

costa ending 6–16 cells before apex, covered and obscured above by chlorophyllose cells, taxifolius-type;

lamina cells irregularly 2-stratose, smooth, bulging, firm-walled, irregularly rounded-hexagonal, 7.5–10 µm. Sexual condition gonioautoicous and cladautoicous;

perigonia gemmiform, axillary, and on short branches;

perichaetia on short axillary branches near proximal ends of stem.

as many as 22 pairs, ligulate to oblong-lanceolate, obtuse-mucronate to acute, to 4 × 0.3–0.5 mm;

dorsal lamina narrowed proximally, ending at insertion or slightly before;

vaginant laminae 1/2–2/3 leaf length, equal in proximal leaves, unequal in distal leaves, minor lamina ending between margin and costa or rounded and ending on or near costa;

margin entire but sometimes serrulate distally, limbate on all laminae, limbidium confluent at apex or ending a few cells before, limbidial cells 2- to 5-stratose;

costa percurrent or ending in mucro, bryoides-type;

laminal cells 1- to irregularly 2-stratose, smooth, slightly bulging, firm-walled, irregularly hexagonal, many somewhat elongate, 10–16 µm, juxtacostal and basal cells of vaginant laminae somewhat larger, quadrate to oblong.

Seta

1.5–5.5 mm.

4–4.5 mm.

Sexual condition

rhizautoicous or synoicous;

perigonia and perichaetia on elongate stems.

Capsule

theca 0.8–2.1 mm, erect, radially symmetric or nearly so;

peristome taxifolius-type;

operculum long rostrate, 0.6–0.9 mm.

theca exserted, erect, radially symmetric to slightly inclined, bilaterally symmetric, 0.6–0.85 mm;

peristome bryoides-type;

operculum 0.4 mm.

Calyptra

cucullate, smooth, 1.5 mm.

not seen.

Spores

13–18 µm.

14–25 µm.

Sporophytes

1 per perichaetium.

1–2 per perichaetium.

Fissidens subbasilaris

Fissidens appalachensis

Habitat Particularly around bases of trees, also rocks, mostly limestone, infrequently decaying wood Crevices and surfaces of rocks submerged in swiftly flowing but usually shallow water, sometimes partially emergent
Distribution
from FNA
AL; AR; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; QC; Mexico (San Luis Potosí, Tamaulipas)
[WildflowerSearch map]
from FNA
NC; PA; SC; TN
Discussion

Fissidens subbasilaris is distinguished, even in the field, by a costa obscured by chlorophyllose cells distally, and short perichaetial branches in the axils of proximal leaves. The plants are typically dark-green and often tinged with red.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens appalachensis, restricted to rapidly moving water, is distinguished by its habitat and strong limbidium. It is most likely to be confused with an expression of F. bryoides that is usually found on wet rocks and stones along the edges of streams, but which differs in its smaller size and weaker limbidium that at times can be partially or completely absent. Laminal cells in size and shape, vaginant laminae, sexuality, thecae, and peristomes in both taxa are similar.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 342. FNA vol. 27, p. 344.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Name authority Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 6–9. (1801) R. H. Zander: Bryologist 72: 406, figs. 1–8. (1969)
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