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fissidens moss, pygmy pocket moss, slender pocket-moss

Habit Plants 1.5–6 × 1 mm. Plants to 2.5 × 2 mm.
Stem

unbranched and branched;

axillary hyaline nodules present;

central strand present.

unbranched;

axillary hyaline nodules absent;

central strand absent.

Leaves

as many as 16 pairs, oblong to lanceolate, acute to short-acuminate, to 1.3 × 0.25 mm;

dorsal lamina narrowed proximally, ending well above or at insertion;

vaginant laminae ± 4/5 leaf length, ± equal, minor lamina ending on or near margin;

margin ± entire to crenulate-serrulate, limbate on all laminae, limbidium variable, developed best on vaginant laminae, ceasing well before apex, limbidium cells 1-stratose;

costa ending 2–5 cells before apex to percurrent, bryoides-type;

laminal cells 1-stratose, distinct, smooth, somewhat bulging, firm-walled, irregularly quadrate to hexagonal, 9–14 µm, increasing somewhat in size and ± oblong in proximal parts of vaginant laminae.

as many as 4 pairs, lanceolate to oblong-lanceolate, acute to obtuse-apiculate, to 2.1 × 0.4 mm;

dorsal lamina ending before insertion;

vaginant laminae 2/5 leaf length, unequal, minor lamina narrowed, ending on or near costa;

margin of dorsal and ventral laminae ± entire to crenate-serrulate, crenate-dentate on vaginant laminae, elimbate;

costa percurrent to ending 2–3 cells before apex, bryoides-type;

laminal cells distinct, smooth, plane, firm-walled, irregularly quadrate to irregularly hexagonal, 10–15 µm, marginal cells slightly smaller, vaginant laminal cells somewhat larger with 2–3 intralaminal rows irregularly elongate.

Seta

2–6 mm.

2–9 mm.

Sexual condition

rhizautoicous;

perigonia gemmiform, proximal to fertile and infertile stems, and as branches along horizontal parts of stems.

rhizautoicous;

perigonia gemmiform, proximal to perichaetial stems or scattered over protonemata.

Capsule

theca exserted, erect, radially symmetric, 0.5–1.25 mm;

peristome anomalous, variable, even in same theca, teeth undivided, imperfectly divided or divided 4/5 their length, papillose distally, smooth proximally;

operculum 0.22–0.3 mm.

theca exserted, erect, radially symmetric, infrequently slightly arcuate, bilaterally symmetric, to 0.7 mm;

operculum 0.4–0.6 mm;

peristome scariosus-type.

Calyptra

cucullate, smooth, to 0.45 mm.

cucullate, smooth, 0.5 mm.

Spores

20–26 µm.

11–14 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens scalaris

Fissidens exilis

Habitat Bare, disturbed soil of stream banks and roadsides, loess banks, often more or less buried Bare, damp soil of shaded banks and in seepage areas
Distribution
from FNA
AL; FL; LA; NC; OK; TX; South America
from FNA
AL; IL; IN; KY; MD; ME; MI; MO; NC; NJ; NY; OH; PA; TN; VT; QC; Europe; Asia; Africa [Introduced in North America; also introduced in West Indies]
Discussion

Fissidens scalaris, although widespread in the southern part of the United States, is infrequently collected. It is best distinguished by its dimorphic stems and undivided to deeply divided, papillose peristome teeth. B. E. Lemmon (1965) published photographs of the development of plants, including perigonial, perichaetial, and infertile stems, and peristome.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens exilis, first reported for North America by W. C. Steere (1950), is probably inadvertently introduced from Europe. It can be distinguished from all other tiny Fissidens in the flora area by the 2–3 rows of irregularly elongate intralaminal cells in the vaginant laminae. Plants of this species would be easily passed over were it not for the persistent protonemata that often form dark green velvetlike carpets over the substrate and the numerous sporophytes that are produced. The gemmiform perigonia are often seen scattered over the protonemata. B. H. Allen et al. (2004) have reviewed and mapped the distribution of F. exilis in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 347. FNA vol. 27, p. 355.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms Moenkemeyera neonii, F. neonii
Name authority Mitten: J. Linn. Soc., Bot. 12: 596. (1869) Hedwig: Sp. Musc. Frond., 152, plate 38, figs. 7–19. (1801)
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