Fissidens scalaris |
Fissidens bushii |
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Bush's fissidens moss |
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Habit | Plants 1.5–6 × 1 mm. | Plants to 13 × 2–2.5 mm. |
Stem | unbranched and branched; axillary hyaline nodules present; central strand present. |
branched; axillary hyaline nodules weak; central strand weak. |
Leaves | as many as 16 pairs, oblong to lanceolate, acute to short-acuminate, to 1.3 × 0.25 mm; dorsal lamina narrowed proximally, ending well above or at insertion; vaginant laminae ± 4/5 leaf length, ± equal, minor lamina ending on or near margin; margin ± entire to crenulate-serrulate, limbate on all laminae, limbidium variable, developed best on vaginant laminae, ceasing well before apex, limbidium cells 1-stratose; costa ending 2–5 cells before apex to percurrent, bryoides-type; laminal cells 1-stratose, distinct, smooth, somewhat bulging, firm-walled, irregularly quadrate to hexagonal, 9–14 µm, increasing somewhat in size and ± oblong in proximal parts of vaginant laminae. |
as many as 20 pairs, oblong to lanceolate, obtuse-apiculate to acute-apiculate, to 2 × 0.5 mm; dorsal lamina rounded proximally, ending at insertion, not decurrent; vaginant laminae ± 2/3 leaf length, ± equal, minor lamina ending on or near margin; margin crenulate-serrulate, more strongly serrulate on vaginant laminae; costa ending 1–2 cells before apex or ending in apiculus, taxifolius-type; laminal cells irregularly 2-stratose in dorsal and ventral laminae, smooth, bulging, firm-walled, rounded-hexagonal, 7–9 µm, papillose in cell corners with 2–4 ± inconspicuous papillae in vaginant laminae. |
Seta | 2–6 mm. |
to 10 mm. |
Sexual condition | rhizautoicous; perigonia gemmiform, proximal to fertile and infertile stems, and as branches along horizontal parts of stems. |
dioicous; perigonia and perichaetia gemmiform, axillary, confined to proximal parts of stems. |
Capsule | theca exserted, erect, radially symmetric, 0.5–1.25 mm; peristome anomalous, variable, even in same theca, teeth undivided, imperfectly divided or divided 4/5 their length, papillose distally, smooth proximally; operculum 0.22–0.3 mm. |
theca to 1 mm, erect, ± arcuate, bilaterally symmetric, about 1 mm; peristome taxifolius-type; operculum 1 mm. |
Calyptra | cucullate, smooth, to 0.45 mm. |
cucullate, smooth, 0.6 mm. |
Spores | 20–26 µm. |
14–18 µm. |
Sporophytes | 1 per perichaetium. |
1 per perichaetium. |
Fissidens scalaris |
Fissidens bushii |
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Habitat | Bare, disturbed soil of stream banks and roadsides, loess banks, often more or less buried | Usually on bare or disturbed clayey soil in open and in woods, along paths, roadside banks, along streams, ravines, infrequently on rocks, stones, tree bases |
Distribution |
AL; FL; LA; NC; OK; TX; South America |
AL; AR; DC; DE; FL; GA; IA; IL; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; VT; WV; NB; NS; ON; QC |
Discussion | Fissidens scalaris, although widespread in the southern part of the United States, is infrequently collected. It is best distinguished by its dimorphic stems and undivided to deeply divided, papillose peristome teeth. B. E. Lemmon (1965) published photographs of the development of plants, including perigonial, perichaetial, and infertile stems, and peristome. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens bushii, named in honor of the collector of the type, Benjamin Franklin Bush, is most apt to be confused with F. taxifolius. However, plants of F. bushii are usually much smaller, costa shorter, and leaves sometimes caducous. Moreover, small papillae restricted to the corners of cells in vaginant laminae are distinctive. The species is probably most closely related to the Asiatic F. teysmannianus Dozy & Molkenboer, which also has inconspicuous papillae in the corners of vaginant laminal cells. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 347. | FNA vol. 27, p. 342. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | Moenkemeyera neonii, F. neonii | F. subbasilaris var. bushii |
Name authority | Mitten: J. Linn. Soc., Bot. 12: 596. (1869) | (Cardot & Thériot) Cardot & Thériot: Bot. Gaz. 37: 365. (1904) |
Web links |