Fissidens fontanus
Fissidens amoenus
fissidens moss, water pocket moss
usually profusely branched;
axillary hyaline nodules absent;
central strand absent.
unbranched;
axillary hyaline nodules present;
central strands absent.
in numerous pairs, lanceolate to linear-lanceolate, frequently somewhat falcate, acute, to 7 × 0.6 mm;
dorsal lamina narrowed proximally, usually ending before insertion;
vaginant laminae 1/3–1/2 the leaf length, unequal, minor lamina usually rounded and free distally or narrowed and ending on or near costa;
margin ± entire, elimbate;
costa ending 15–35 cells before apex, bryoides-type;
laminal cells usually 1-stratose, sometimes 2-stratose juxtacostally, quadrate to short-oblong to hexagonal, smooth, slightly bulging, ± firm-walled, 15–23 × 10–18 µm in dorsal and ventral laminae, juxtacostal cells largest.
as many as 10 pairs, ligulate to lanceolate, acute, usually apiculate, to 1.8 × 0.25 mm;
dorsal lamina narrowed proximally, ending at insertion, not decurrent;
vaginant laminae ± 1/2 leaf length, acute, ± unequal, minor lamina ending near margin;
margin ± evenly crenulate-serrulate, elimbate except for weak limbidium on proximal parts of vaginant laminae of perichaetial leaves;
costa percurrent to ending as many as 5 cells before apex, bryoides-type;
laminal cells 1-stratose, distinct, smooth, often strongly bulging, firm-walled, irregularly quadrate to hexagonal, 8–11 µm, larger and ± oblong juxtacostally in proximal parts of vaginant laminae.
0.5–0.6 mm.
to 5 mm.
gonioautoicous and cladautoicous;
perigonia and perichaetia on short axillary branches, 1–5 per leaf axil, sometimes on elongate, axillary branches.
rhizautoicous and gonioautoicous.
theca 0.5–0.6 mm, erect, radially symmetric;
peristome anomalous, irregularly divided, ± incomplete, often truncate, papillose;
operculum as long as seta.
theca exserted, erect, radially symmetric, to 0.6 mm;
peristome bryoides-type;
operculum 0.5 mm.
mitrate, smooth.
cucullate, smooth, 0.5 mm.
18–27 µm.
11–16 µm.
1–5 per leaf axil, inconspicuous.
1 per perichaetium.
Fissidens fontanus
Fissidens amoenus
Fissidens fontanus and F. hallianus are aquatic species usually found in stagnant or slow moving water, but at times emergent because of fluctuating water levels (H. A. Crum and L. E. Anderson 1981; P. G. Davison and D. E. Wujek 1999). Plants of both species are usually profusely branched, delicate, and feathery in appearance with leaves often 10 times or more as long as wide. Often plants are encrusted with diatoms. Stems and leaves might also appear black due to deposition of organic matter. When dry the leaves are unusually fragile so that when being prepared for herbarium storage it is best to float plants onto sheets prior to drying.
A suite of gametophytic and sporophytic characters distinguishes Fissidens fontanus. The unequal vaginant laminae are about 1/4–1/3 the leaf length, and minor lamina is round and more or less free distally. In addition, costa ends well below the leaf apex. In fertile specimens, the clusters of 1–5 axillary, inconspicuous, dark sporophytes in each perichaetium are distinctive. The sporophytes, more prevalent than thought, are caducous and are often found in the bottom of the herbarium packets. The short seta is equal in length to that of operculum, and peristome teeth are greatly reduced, often consisting of only the truncated, undivided basal parts.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
A small, attractive species, Fissidens amoenus can be recognized by its distinct laminal cells, crenulate-serrulate leaf margin, large, more or less oblong, pellucid cells in the proximal parts of the vaginant laminae, and weak limbidium restricted to the proximal parts of the vaginant laminae of perichaetial leaves. The laminal cells have been described as “inconspicuously 1-papillose” (H. A. Crum and L. E. Anderson 1981, as F. hallii), but since that condition is not always present it is perhaps best to regard these cells as strongly bulging.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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