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elegant fissidens moss

Habit Plants to 5.5 × 2 mm. Plants 1.5–6 × 1 mm.
Stem

unbranched and branched;

axillary hyaline nodules absent;

central strand weak.

unbranched and branched;

axillary hyaline nodules present;

central strand present.

Leaves

as many as 12 pairs, oblong to lanceolate, acute to obtuse-apiculate, most ending in a clear, sharp cell, to 1.8 × 0.4 mm;

dorsal lamina narrowed or rounded proximally, ending at or before insertion, not decurrent;

vaginant laminae 1/2–2/3 leaf length, unequal, minor lamina ending near margin;

margin serrulate, limbate and entire to remotely denticulate on proximal 2/3 or less of vaginant laminae of most leaves of perichaetial stems, often absent from leaves of infertile stems, limbidium intralaminal in part or completely, limbidial cells 1-stratose;

costa ending 2–4 cells before apex or percurrent, ending in apiculus, infrequently short-excurrent, bryoides-type;

lamina cells 1-stratose, pluripapillose, obscure, firm-walled, rounded-hexagonal, 5–8 µm, twice as deep as wide.

as many as 16 pairs, oblong to lanceolate, acute to short-acuminate, to 1.3 × 0.25 mm;

dorsal lamina narrowed proximally, ending well above or at insertion;

vaginant laminae ± 4/5 leaf length, ± equal, minor lamina ending on or near margin;

margin ± entire to crenulate-serrulate, limbate on all laminae, limbidium variable, developed best on vaginant laminae, ceasing well before apex, limbidium cells 1-stratose;

costa ending 2–5 cells before apex to percurrent, bryoides-type;

laminal cells 1-stratose, distinct, smooth, somewhat bulging, firm-walled, irregularly quadrate to hexagonal, 9–14 µm, increasing somewhat in size and ± oblong in proximal parts of vaginant laminae.

Seta

to 5 mm.

2–6 mm.

Sexual condition

rhizautoicous and gonioautoicous.

rhizautoicous;

perigonia gemmiform, proximal to fertile and infertile stems, and as branches along horizontal parts of stems.

Capsule

theca exserted, erect, radially symmetric, to 0.8 mm;

peristome scariosus-type;

operculum to 0.5 mm.

theca exserted, erect, radially symmetric, 0.5–1.25 mm;

peristome anomalous, variable, even in same theca, teeth undivided, imperfectly divided or divided 4/5 their length, papillose distally, smooth proximally;

operculum 0.22–0.3 mm.

Calyptra

cucullate, smooth, to 0.6 mm.

cucullate, smooth, to 0.45 mm.

Spores

9–13 µm.

20–26 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens elegans

Fissidens scalaris

Habitat Sandy and clayey soils along roadsides and streams, banks of ravines, bluffs, loess banks, uprooted tree roots, trunks of trees, decaying stumps, sandstone Bare, disturbed soil of stream banks and roadsides, loess banks, often more or less buried
Distribution
from FNA
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; SC; TN; TX; VA; Mexico; Central America; South America; West Indies
[WildflowerSearch map]
from FNA
AL; FL; LA; NC; OK; TX; South America
Discussion

The obscure, pluripapillose lamina cells and leaf apex that terminates in a single, hyaline, sharply pointed cell distinguish Fissidens elegans. The limbidium is quite variable, occurring on the vaginant laminae of most leaves of perichaetial stems to being absent from the leaves of infertile stems. The species is close to F. pallidinervis (see discussion under 26). H. A. Crum and L.E. Anderson (1981) commented on a collection that was possibly made in Lafayette, Wisconsin, a site quite distant from the normal distribution of this species in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens scalaris, although widespread in the southern part of the United States, is infrequently collected. It is best distinguished by its dimorphic stems and undivided to deeply divided, papillose peristome teeth. B. E. Lemmon (1965) published photographs of the development of plants, including perigonial, perichaetial, and infertile stems, and peristome.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 350. FNA vol. 27, p. 347.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. ravenelii Moenkemeyera neonii, F. neonii
Name authority Bridel: Muscol. Recent., suppl. 1: 167. (1806) Mitten: J. Linn. Soc., Bot. 12: 596. (1869)
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