Fissidens elegans |
Fissidens adianthoides |
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elegant fissidens moss |
fissidens moss, maidenhair pocket-moss |
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Habit | Plants to 5.5 × 2 mm. | Plants to 85 × 5 mm. |
Stem | unbranched and branched; axillary hyaline nodules absent; central strand weak. |
mostly branched; axillary hyaline nodules absent; central strand present. |
Leaves | as many as 12 pairs, oblong to lanceolate, acute to obtuse-apiculate, most ending in a clear, sharp cell, to 1.8 × 0.4 mm; dorsal lamina narrowed or rounded proximally, ending at or before insertion, not decurrent; vaginant laminae 1/2–2/3 leaf length, unequal, minor lamina ending near margin; margin serrulate, limbate and entire to remotely denticulate on proximal 2/3 or less of vaginant laminae of most leaves of perichaetial stems, often absent from leaves of infertile stems, limbidium intralaminal in part or completely, limbidial cells 1-stratose; costa ending 2–4 cells before apex or percurrent, ending in apiculus, infrequently short-excurrent, bryoides-type; lamina cells 1-stratose, pluripapillose, obscure, firm-walled, rounded-hexagonal, 5–8 µm, twice as deep as wide. |
as many as 60 pairs, somewhat undulate, oblong to lanceolate, acute, occasionally obtuse, to 3.5 × 1.2 mm; dorsal lamina rounded or narrowed proximally, ending at insertion, not decurrent; vaginant laminae 1/2–3/5 the leaf length, ± equal, minor lamina ending on or near margin; margin crenulate to regularly serrulate but irregularly serrate distally, elimbate; costa ending 2–3 cells before apex to percurrent, taxifolius-type; laminal cells usually 1-stratose, infrequently and irregularly 2-stratose in the dorsal and ventral laminae, distinct, smooth, ± plane, firm-walled, irregularly quadrate to hexagonal, 10–20 µm, 1–5 marginal rows usually thinner with slightly thicker walls, forming a lighter marginal band, juxtacostal cells larger, mostly oblong and pellucid; vaginant laminal cells somewhat larger. |
Seta | to 5 mm. |
to 25 mm. |
Sexual condition | rhizautoicous and gonioautoicous. |
dioicous (?); perigonia not seen; perichaetia on short axillary branches, generally confined to proximal half of stems. |
Capsule | theca exserted, erect, radially symmetric, to 0.8 mm; peristome scariosus-type; operculum to 0.5 mm. |
theca mostly ± inclined, ± arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum to 1.5 mm. |
Calyptra | cucullate, smooth, to 0.6 mm. |
cucullate, smooth, to 2.5 mm. |
Spores | 9–13 µm. |
13–22 µm. |
Sporophyte(s) | 1 per perichaetium. |
1 per perichetium. |
Fissidens elegans |
Fissidens adianthoides |
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Habitat | Sandy and clayey soils along roadsides and streams, banks of ravines, bluffs, loess banks, uprooted tree roots, trunks of trees, decaying stumps, sandstone | Seepage areas, along streams, near waterfalls, meadows, soil, around bases of trees, decaying wood, dripping limestone and sandstone rocks and boulders |
Distribution |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; SC; TN; TX; VA; Mexico; Central America; South America; West Indies
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AK; AL; AR; CA; CT; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NJ; NY; OH; OK; OR; PA; RI; TN; TX; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; NT; NU; ON; QC; SK; YT; Asia; Greenland; Europe; Africa
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Discussion | The obscure, pluripapillose lamina cells and leaf apex that terminates in a single, hyaline, sharply pointed cell distinguish Fissidens elegans. The limbidium is quite variable, occurring on the vaginant laminae of most leaves of perichaetial stems to being absent from the leaves of infertile stems. The species is close to F. pallidinervis (see discussion under 26). H. A. Crum and L.E. Anderson (1981) commented on a collection that was possibly made in Lafayette, Wisconsin, a site quite distant from the normal distribution of this species in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The usually 1-stratose, distinct, smooth, plane laminal cells, lighter band of marginal laminal cells, and short perichaetial stems in the axils of proximal leaves distinguish Fissidens adianthoides. The species, nevertheless, can be confused with F. osmundioides, which has similar laminal cells without a marginal band of lighter cells and a serrate leaf apex, but which differs in having terminal perichaetia and papillose rhizoids. Fissidens dubius, which also has short perichaetial stems in the axils of proximal leaves and often a lighter band of marginal laminal cells, differs from F. adianthoides in its smaller and rather obscure laminal cells that are consistently irregularly 2-stratose. Fissidens adianthoides and F. dubius (as F. cristatus), according to cytological evidence presented by L. E. Anderson and V. S. Bryan (1956), are distantly related. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 350. | FNA vol. 27, p. 339. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. ravenelii | F. adianthoides var. immarginatus |
Name authority | Bridel: Muscol. Recent., suppl. 1: 167. (1806) | Hedwig: Sp. Musc. Frond., 157. (1801) |
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