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elegant fissidens moss

pale-nerve fissidens moss

Habit Plants to 5.5 × 2 mm. Plants to 6 × 1 mm.
Stem

unbranched and branched;

axillary hyaline nodules absent;

central strand weak.

branched and unbranched;

axillary hyaline nodules absent;

central strand weak or absent.

Leaves

as many as 12 pairs, oblong to lanceolate, acute to obtuse-apiculate, most ending in a clear, sharp cell, to 1.8 × 0.4 mm;

dorsal lamina narrowed or rounded proximally, ending at or before insertion, not decurrent;

vaginant laminae 1/2–2/3 leaf length, unequal, minor lamina ending near margin;

margin serrulate, limbate and entire to remotely denticulate on proximal 2/3 or less of vaginant laminae of most leaves of perichaetial stems, often absent from leaves of infertile stems, limbidium intralaminal in part or completely, limbidial cells 1-stratose;

costa ending 2–4 cells before apex or percurrent, ending in apiculus, infrequently short-excurrent, bryoides-type;

lamina cells 1-stratose, pluripapillose, obscure, firm-walled, rounded-hexagonal, 5–8 µm, twice as deep as wide.

as many as 18 pairs, lingulate to lanceolate, rounded to obtuse to broadly acute, to 1 × 0.25 mm;

dorsal lamina narrowed or rounded proximally, ending at insertion, not decurrent;

vaginant laminae ± 1/2–2/3 leaf length, ± unequal, minor lamina ending near margin;

margin serrulate, limbate, ± entire in proximal 1/3–1/2 of perichaetial and subtending 1–2 pairs of leaves, limbidium sometimes indistinct, limbidial cells 1-stratose;

costa ending 3–25 cells before apex, often spurred distally, bryoides-type;

laminal cells 1-stratose, pluripapillose, obscure, firm-walled, irregularly quadrate to hexagonal, 4–8 µm, in transverse section usually twice as deep as wide.

Seta

to 5 mm.

to 2 mm.

Sexual condition

rhizautoicous and gonioautoicous.

rhizautoicous, cladautoicous, rarely gonioautoicous.

Capsule

theca exserted, erect, radially symmetric, to 0.8 mm;

peristome scariosus-type;

operculum to 0.5 mm.

theca exserted, erect, radially symmetric, to 0.6 mm;

peristome scariosus-type;

operculum 0.3 mm.

Calyptra

cucullate, smooth, to 0.6 mm.

cucullate, ± prorate, 0.3 µm. Spores 9–14µm.

Spores

9–13 µm.

Sporophytes

1 per perichaetium.

1–2 per perichaetium.

Fissidens elegans

Fissidens pallidinervis

Habitat Sandy and clayey soils along roadsides and streams, banks of ravines, bluffs, loess banks, uprooted tree roots, trunks of trees, decaying stumps, sandstone Usually in damp areas around bases of trees, decaying logs, occasionally limestone and soil
Distribution
from FNA
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; SC; TN; TX; VA; Mexico; Central America; South America; West Indies
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from FNA
FL; LA; Mexico; Central America; South America; West Indies; Asia; Africa
Discussion

The obscure, pluripapillose lamina cells and leaf apex that terminates in a single, hyaline, sharply pointed cell distinguish Fissidens elegans. The limbidium is quite variable, occurring on the vaginant laminae of most leaves of perichaetial stems to being absent from the leaves of infertile stems. The species is close to F. pallidinervis (see discussion under 26). H. A. Crum and L.E. Anderson (1981) commented on a collection that was possibly made in Lafayette, Wisconsin, a site quite distant from the normal distribution of this species in North America.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens pallidinervis and F. elegans are the only species of the genus in North America with small, obscure, pluripapillose laminal cells. Fissidens pallidinervis, however, is distinguished by a rounded to broadly acute leaf apex, and a limbidium restricted to the lower parts of vaginant laminae of perichaetial and one or two pairs of subtending leaves. Axillary, stalked, multicellular, clavate gemmae have been reported in Japanese collections of F. pallidinervis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 350. FNA vol. 27, p. 350.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. ravenelii F. garberi, F. minutus
Name authority Bridel: Muscol. Recent., suppl. 1: 167. (1806) Mitten: J. Linn. Soc., Bot. 12: 592. (1869)
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