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fissidens moss

Habit Plants to 25 × 3.5 mm. Plants to 7 × 2.5 mm.
Stem

usually branched;

axillary hyaline nodules absent;

central strand present.

unbranched and branched;

axillary hyaline nodules absent;

central strand absent.

Leaves

as many as 25 pairs, often undulate, oblong to lanceolate, obtuse to acute, often apiculate, to 3.5 × 0.7 mm;

dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent;

vaginant laminae 2/3 the leaf length, ± unequal, minor lamina ending near margin;

margin crenulate to crenulate-serrulate but unevenly serrate at leaf apex, elimbate;

costa ending a few cells before apex to percurrent, taxifolius-type;

laminal cells irregularly 2-stratose in dorsal and ventral laminae, ± obscure, smooth, bulging, firm-walled, rounded-hexagonal, 7–10 µm, 3–4 rows of marginal cells thinner, walls slightly thicker, forming a pale marginal band.

as many as 12 pairs, oblong to lingulate, broadly acute to obtuse, sometimes apiculate, to 1.4 × 0.5 mm;

dorsal lamina narrowed proximally, ending at insertion, not decurrent;

vaginant laminae ± 1/2 leaf length, ± unequal, minor lamina ending near margin;

margin crenulate-serrulate but limbate and entire on proximal 1/3–1/2 of vaginant laminae of most leaves, limbidial cells 1-stratose;

costa ending 2–6 cells before apex, bryoides-type;

lamina cells 1-stratose, distinct, mammillose, firm-walled, irregularly hexagonal, 7–12 µm, smooth, larger, ± oblong, pellucid juxtacostally in proximal parts of vaginant laminae.

Seta

to 11 mm.

to 2 mm.

Sexual condition

dioicous;

perigonia on short axillary branches and elongate stems;

perichaetia on short axillary branches, mostly near proximal ends of stems.

gonioautoicous.

Capsule

theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.8 mm;

peristome taxifolius-type;

operculum 1–1.4 mm.

theca exserted, erect, radially symmetric, 0.5 mm;

peristome scariosus-type;

operculum 0.4 mm.

Calyptra

cucullate, smooth, 1.8 mm.

cucullate, smooth, 0.3–0.5 mm.

Spores

13–20 µm.

10–16 µm.

Sporophytes

1 per perchaetium.

1 per perichaetium.

Fissidens dubius

Fissidens leptophyllus

Habitat Soil and humus, bases of trees, decaying wood, rocks and boulders in moist, shaded sites Around bases of trees, decaying wood, infrequently on soil, cypress-gum swamps
Distribution
from FNA
AK; AL; AR; AZ; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; MA; ME; MI; MN; MO; MS; NC; NE; NH; NY; OK; PA; SC; TN; TX; VA; VT; WI; WV; NB; NF; NS; ON; QC; Mexico; Central America; West Indies; Europe; Asia; Pacific Islands
[WildflowerSearch map]
from FNA
FL; LA
Discussion

In contrast to Fissidens adianthoides, F. dubius has smaller, irregularly 2-stratose, obscure laminal cells. Subterranean, irregularly globose, multicellular rhizoidal gemmae have been reported for F. dubius as well as F. osmundioides and F. taxifolius in Europe and Japan. Z. Iwatsuki and T. Suzuki (1982) cited the occurrence of dwarf male plants (small perigonial stems located on the leaves of perichaetial plants) in Japanese specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The extent of the limbidium is the best way to distinguish Fissidens leptophyllus from F. submarginatus. The limbidium in F. leptophyllus is confined to the proximal 1/2 or less of the vaginant laminae of most leaves, but in F. submarginatus the limbidium extends the entire length of the vaginant laminae and sometimes onto the proximal part of the adjacent ventral lamina of most or all leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 340. FNA vol. 27, p. 353.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. adianthoides var. semicristatus, F. cristatus, F. cristatus var. winonensis, F. decipiens var. winonensis, F. floridanus F. reesei
Name authority P. Beauvois: Prodr. Aethéogam., 57. (1805) Montagne: Ann. Sci. Nat., Bot., sér. 2, 14: 344. (1840)
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