Fissidens curvatus |
Fissidens submarginatus |
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Habit | Plants 1.2–10 × 1–2.5 mm. | Plants to 5 × 2 mm. |
Stem | unbranched and branched; axillary hyaline nodules present; central strand weak or absent. |
unbranched and branched; axillary hyaline nodules absent; central strand weak. |
Leaves | as many as 25 pairs, narrowly lanceolate to ovate–lanceolate, acute to sharply acute to short acuminate, to 1.9 × 0.3; dorsal lamina narrowed proximally, ending before or at insertion, infrequently ± decurrent; vaginant laminae 1/2–4/5 leaf length, ± equal, minor lamina ending on or near margin; margin entire, limbate on all laminae, limbidium confluent at apex or ending shortly before apex, extending to base of dorsal lamina, limbidial cells 2–3-stratose; costa percurrent to excurrent, bryoides-type; laminal cells 1-stratose, distinct, smooth, ± bulging, firm-walled, elongate, 7–22 × 4–13 µm, increasing in size toward proximal part of leaf. |
as many as 10 pairs, lanceolate to lingulate, acute to obtuse-apiculate, to 1.3 × 0.4 mm; dorsal lamina narrowed abruptly proximally, ending at insertion, not decurrent; vaginant laminae ± 1/2 leaf length, equal; margin serrulate, limbate and entire on complete length of vaginant laminae and often on proximal portion of adjacent ventral laminae of all leaves except smallest, limbidial cells 1-stratose; costa percurrent to ending 2–3 cells before apex, bryoides-type; laminal cells 1-stratose, distinct, mammillose, firm-walled, quadrate to hexagonal, 7–9 µm. |
Seta | to 12 mm. |
to 3.5 mm. |
Sexual condition | rhizoautoicous; perigonia gemmiform, proximal to infertile and fertile stems. |
rhizautoicous; perigonia gemmiform, proximal to fertile and infertile stems. |
Capsule | theca exserted, ± erect, radially symmetric to inclined, arcuate, bilaterally symmetic, 0.35–1.25 mm; peristome bryoides-type; operculum 0.25–0.3 µm. |
theca exserted, erect, radially symmetric, to 0.6 mm; peristome scariosus-type; operculum to 9.4 mm. |
Calyptra | cucullate, smooth, 0.5 mm. |
cucullate, smooth, to 0.5 mm. |
Spores | 11–18 µm. |
11–23 µm. |
Sporophytes | 1 per perichaetium. |
1 per perichaetium. |
Fissidens curvatus |
Fissidens submarginatus |
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Habitat | Bare, shaded soil, often among grasses in open Quercus forests and around bases of coastal shrubs | Moist, disturbed soil and around bases of trees, mostly in cypress swamps |
Distribution |
CA; Mexico; South America; West Indies; Europe; Asia; Africa; Pacific Islands (New Caledonia, New Zealand); Australia |
LA; Mexico; Central America; South America; West Indies; Africa |
Discussion | Fissidens curvatus is distinguished by its delicate dimorphic stems, usually long excurrent costa, limbidium confluent at leaf apex, and slightly elongate laminal cells. The theca is usually arcuate but can also be erect. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens submarginatus is most likely to be confused with F. leptophyllus. Both species have mammillose laminal cells, serrulate leaf margin, and limbidium of 1-stratose cells restricted essentially to vaginant laminae. The limbidium is the best character by which these two species are separated. In F. submarginatus the limbidium consistently extends the entire length of the vaginant laminae of most or all leaves, and on occasion extends a short distance onto the proximal part of the adjacent ventral lamina. In contrast, the limbidium in F. leptophyllus consistently extends no more than 1/2 the length of the vaginant laminae of most leaves. Minor differences can also be found in the shape of the leaf apex (usually apiculate vs. not apiculate), costa length (ending 2–3 cells before the leaf apex vs. ending 2–6 cells before the leaf apex), and cell size (7–9 µm versus 7–12 µm). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 347. | FNA vol. 27, p. 352. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. milobakeri | F. kochii |
Name authority | Hornschuch: Linnaea 15: 148. (1841) | Bruch: Flora 29: 133. (1846) |
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