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polypody fissidens moss

Habit Plants 1.2–10 × 1–2.5 mm. Plants to 50 × 6.5 mm.
Stem

unbranched and branched;

axillary hyaline nodules present;

central strand weak or absent.

unbranched and sparingly branched;

axillary hyaline nodules absent;

central strand present.

Leaves

as many as 25 pairs, narrowly lanceolate to ovate–lanceolate, acute to sharply acute to short acuminate, to 1.9 × 0.3;

dorsal lamina narrowed proximally, ending before or at insertion, infrequently ± decurrent;

vaginant laminae 1/2–4/5 leaf length, ± equal, minor lamina ending on or near margin;

margin entire, limbate on all laminae, limbidium confluent at apex or ending shortly before apex, extending to base of dorsal lamina, limbidial cells 2–3-stratose;

costa percurrent to excurrent, bryoides-type;

laminal cells 1-stratose, distinct, smooth, ± bulging, firm-walled, elongate, 7–22 × 4–13 µm, increasing in size toward proximal part of leaf.

as many as 80 pairs, somewhat undulate, oblong, obtuse to broadly acute, usually bluntly mucronate, to 4 × 1 mm;

dorsal lamina rounded proximally, ending at insertion;

vaginant laminae 1/2 the leaf length, acute, ± equal, minor lamina ending on or near margin;

margin entire but denticulate distally, elimbate;

costa percurrent to ending 2–4 cells before apex, taxifolius-type;

laminal cells 1-stratose, distinct, smooth, slightly bulging, firm-walled, irregularly hexagonal, 10–20 µm, larger juxtacostally, smaller at margin.

Seta

to 12 mm.

to 15 mm.

Sexual condition

rhizoautoicous;

perigonia gemmiform, proximal to infertile and fertile stems.

dioicous;

perigonia and perichaetia on short axillary branches and main stems.

Capsule

theca exserted, ± erect, radially symmetric to inclined, arcuate, bilaterally symmetic, 0.35–1.25 mm;

peristome bryoides-type;

operculum 0.25–0.3 µm. Calyptra cucullate, smooth, 0.5 mm.

theca slightly inclined, radially symmetric or slightly arcuate, bilaterally symmetric, to 2.5 mm;

operculum about as long as theca.

Calyptra

cucullate, smooth, 1.5–1.55 mm.

Spores

11–18 µm.

9–11 µm.

Sporophytes

1 per perichaetium.

1–2 per perichaetium.

Fissidens curvatus

Fissidens polypodioides

Habitat Bare, shaded soil, often among grasses in open Quercus forests and around bases of coastal shrubs Calcareous soil, and limestone rocks and boulders along streams and in ravines
Distribution
from FNA
CA; Mexico; South America; West Indies; Europe; Asia; Africa; Pacific Islands (New Caledonia, New Zealand); Australia
from FNA
AL; FL; GA; IN; LA; MS; NC; SC; TX; Mexico; Central America; South America; West Indies; Asia
Discussion

Fissidens curvatus is distinguished by its delicate dimorphic stems, usually long excurrent costa, limbidium confluent at leaf apex, and slightly elongate laminal cells. The theca is usually arcuate but can also be erect.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens polypodioides is a robust species and has its greatest distribution in the neotropics and Asia. It is perhaps the handsomest species of the genus in the flora area. However, it might be confused with F. asplenioides, from which it is distinguished by taxifolius-type costa, leaves crisped but not curled tightly inward when dry, entire leaf margin, acute and more or less equal vaginant laminae, larger laminal cells that are not lenticularly thickened, and absent differentiated medial marginal cells in the vaginant laminae. The central strand is especially well developed. H. A. Crum and L. E. Anderson (1965, 1981) published illustrations of F. polypodioides and F. asplenioides that aid greatly in distinguishing the two.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 347. FNA vol. 27, p. 339.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. milobakeri
Name authority Hornschuch: Linnaea 15: 148. (1841) Hedwig: Sp. Musc. Frond., 154. (1801)
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