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fissidens moss, poor pocket moss

Habit Plants 1.2–10 × 1–2.5 mm. Plants 2–3 × 1.5–2.5 mm.
Stem(s)

unbranched and branched;

axillary hyaline nodules present;

central strand weak or absent.

unbranched;

axillary hyaline nodules absent;

central strands weak.

Leaves

as many as 25 pairs, narrowly lanceolate to ovate–lanceolate, acute to sharply acute to short acuminate, to 1.9 × 0.3;

dorsal lamina narrowed proximally, ending before or at insertion, infrequently ± decurrent;

vaginant laminae 1/2–4/5 leaf length, ± equal, minor lamina ending on or near margin;

margin entire, limbate on all laminae, limbidium confluent at apex or ending shortly before apex, extending to base of dorsal lamina, limbidial cells 2–3-stratose;

costa percurrent to excurrent, bryoides-type;

laminal cells 1-stratose, distinct, smooth, ± bulging, firm-walled, elongate, 7–22 × 4–13 µm, increasing in size toward proximal part of leaf.

as many as 5 pairs, oblong to oblanceolate to lanceolate, acute to short-acuminate, to 2.1 × 0.6 mm;

margin crenulate-serrulate but often sharply serrate on distal parts of vaginant laminae, elimbate;

dorsal lamina usually narrowed proximally, ending at insertion, not decurrent;

vaginant laminae ± 1/2–2/3 leaf length, unequal, minor lamina ending near margin;

costa ending 6–15 cells before apex, bryoides-type;

laminal cells 1-stratose, distinct, smooth, plane, firm-walled, long-hexagonal to oblong to regularly quadrate, decreasing in size from costa to margin, medial cells 10–54 × 12–25 µm, largest along costa, marginal cells 9–15 µm, juxtacostal cells in vaginant laminae pellucid, greatly enlarged.

Seta

to 12 mm.

2.5–5 mm.

Sexual condition

rhizoautoicous;

perigonia gemmiform, proximal to infertile and fertile stems.

rhizautoicous and gonioautoicous.

Capsule

theca exserted, ± erect, radially symmetric to inclined, arcuate, bilaterally symmetic, 0.35–1.25 mm;

peristome bryoides-type;

operculum 0.25–0.3 µm.

theca exserted, usually inclined, slightly arcuate, bilaterally symmetric, infrequently erect, nearly radially symmetric, 0.7 mm;

peristome scariosus-type;

operculum 0.3–0.8 mm.

Calyptra

cucullate, smooth, 0.5 mm.

cucullate, smooth, 0.4 mm.

Spores

11–18 µm.

9–13 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens curvatus

Fissidens pauperculus

Habitat Bare, shaded soil, often among grasses in open Quercus forests and around bases of coastal shrubs Bare gravelly soil in dried stream beds and on banks, sometimes found with F. crispus, often associated with the Coast Redwood Forest (L. F. Koch 1951)
Distribution
from FNA
CA; Mexico; South America; West Indies; Europe; Asia; Africa; Pacific Islands (New Caledonia, New Zealand); Australia
from FNA
CA; OR; WA; BC
Discussion

Fissidens curvatus is distinguished by its delicate dimorphic stems, usually long excurrent costa, limbidium confluent at leaf apex, and slightly elongate laminal cells. The theca is usually arcuate but can also be erect.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens pauperculus is distinguished by its short costa, and smooth, plane laminal cells that gradually decrease in size outwardly. P. A. Florschütz (1964) equated specimens from Suriname with this species. R. A. Pursell (1994b) demonstrated, however, that the North American plants are different from those of South America. Although close to the South American plants that belong to F. inaequalis Mitten, the North American species differs by its slightly smaller laminal cells and in the absence of truncate-retuse vaginant laminae in the perichaetial leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 347. FNA vol. 27, p. 354.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. milobakeri
Name authority Hornschuch: Linnaea 15: 148. (1841) Howe: Erythea 2: 97, fig. 1. (1894)
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