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fissidens moss

Habit Plants 1.2–10 × 1–2.5 mm. Plants to 120 × 6 mm.
Stem

unbranched and branched;

axillary hyaline nodules present;

central strand weak or absent.

usually profusely branched;

axillary hyaline nodules absent;

central strand absent.

Leaves

as many as 25 pairs, narrowly lanceolate to ovate–lanceolate, acute to sharply acute to short acuminate, to 1.9 × 0.3;

dorsal lamina narrowed proximally, ending before or at insertion, infrequently ± decurrent;

vaginant laminae 1/2–4/5 leaf length, ± equal, minor lamina ending on or near margin;

margin entire, limbate on all laminae, limbidium confluent at apex or ending shortly before apex, extending to base of dorsal lamina, limbidial cells 2–3-stratose;

costa percurrent to excurrent, bryoides-type;

laminal cells 1-stratose, distinct, smooth, ± bulging, firm-walled, elongate, 7–22 × 4–13 µm, increasing in size toward proximal part of leaf.

in numerous pairs, lanceolate to linear-lanceolate, acute, to 6.5 × 0.7 mm;

dorsal lamina narrowed proximally, usually ending before insertion;

vaginant laminae 1/4–1/2 leaf length, acute, unequal in most leaves, minor lamina rounded above, attached on or near costa, ± equal on proximal part of stem, minor lamina ending near leaf margin;

margin ± entire, elimbate or limbate in the proximal 2/3 or less of vaginant laminae;

costa ending 5–15 cells before apex, bryoides-type;

laminal cells 1-stratose, distinct, smooth, slightly bulging, ± firm-walled, quadrate to oblong to hexagonal, 10–15 × 9–11 µm.

Seta

to 12 mm.

0.7–1.5 mm, often bent at the vaginula.

Sexual condition

rhizoautoicous;

perigonia gemmiform, proximal to infertile and fertile stems.

cladautoicous;

perigonia on elongate, axillary branches;

perichaetia terminal on mains stems and elongate, axillary branches.

Capsule

theca exserted, ± erect, radially symmetric to inclined, arcuate, bilaterally symmetic, 0.35–1.25 mm;

peristome bryoides-type;

operculum 0.25–0.3 µm.

theca erect, radially symmetric, 0.4–0.8 mm;

peristome anomalous, teeth variable, undivided and often fenestrate to divided 2/3 their length, even in same capsule, nearly smooth to papillose proximally, filaments with straight or spiral thickenings or striations;

operculum 0.4–0.6 mm.

Calyptra

cucullate, smooth, 0.5 mm.

not seen.

Spores

11–18 µm.

14–32 µm.

Sporophytes

1 per perichaetium.

1–2 per perichaetium.

Fissidens curvatus

Fissidens hallianus

Habitat Bare, shaded soil, often among grasses in open Quercus forests and around bases of coastal shrubs Flowing and non-flowing water over limestone, granite, and bases of trees, especially abundant in Taxodium-Nyssa swamps
Distribution
from FNA
CA; Mexico; South America; West Indies; Europe; Asia; Africa; Pacific Islands (New Caledonia, New Zealand); Australia
from FNA
FL; IL; LA; MA; MS; NC; NJ; TX
Discussion

Fissidens curvatus is distinguished by its delicate dimorphic stems, usually long excurrent costa, limbidium confluent at leaf apex, and slightly elongate laminal cells. The theca is usually arcuate but can also be erect.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens hallianus, named for Elihu Hall, has been confused with F. fontanus; they can be found growing together and both have similar vaginant laminae, although in F. hallianus there can be a weak limbidiuim on the proximal margin. Moreover, the costa is longer in F. hallianus than in F. fontanus. The sporophytes of F. hallianus are terminal, the number per perichaetium usually fewer than in F. fontanus, the seta is longer, relative to the length of the capsule, and the peristome teeth are complete. Sexual reproduction appears to be dependent on seasonal exposure.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 347. FNA vol. 27, p. 344.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. milobakeri Conomitrium hallianum, F. manateensis, Octodiceras hallianum
Name authority Hornschuch: Linnaea 15: 148. (1841) (Sullivant & Lesquereux) Mitten: J. Linn. Soc., Bot. 21: 560. (1885)
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