Fissidens curvatus |
Fissidens appalachensis |
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Appalachian fissidens moss |
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Habit | Plants 1.2–10 × 1–2.5 mm. | Plants to 13 × 2–4 mm. |
Stem | unbranched and branched; axillary hyaline nodules present; central strand weak or absent. |
unbranched and branched; axillary hyaline nodules weak or absent; central strand weak. |
Leaves | as many as 25 pairs, narrowly lanceolate to ovate–lanceolate, acute to sharply acute to short acuminate, to 1.9 × 0.3; dorsal lamina narrowed proximally, ending before or at insertion, infrequently ± decurrent; vaginant laminae 1/2–4/5 leaf length, ± equal, minor lamina ending on or near margin; margin entire, limbate on all laminae, limbidium confluent at apex or ending shortly before apex, extending to base of dorsal lamina, limbidial cells 2–3-stratose; costa percurrent to excurrent, bryoides-type; laminal cells 1-stratose, distinct, smooth, ± bulging, firm-walled, elongate, 7–22 × 4–13 µm, increasing in size toward proximal part of leaf. |
as many as 22 pairs, ligulate to oblong-lanceolate, obtuse-mucronate to acute, to 4 × 0.3–0.5 mm; dorsal lamina narrowed proximally, ending at insertion or slightly before; vaginant laminae 1/2–2/3 leaf length, equal in proximal leaves, unequal in distal leaves, minor lamina ending between margin and costa or rounded and ending on or near costa; margin entire but sometimes serrulate distally, limbate on all laminae, limbidium confluent at apex or ending a few cells before, limbidial cells 2- to 5-stratose; costa percurrent or ending in mucro, bryoides-type; laminal cells 1- to irregularly 2-stratose, smooth, slightly bulging, firm-walled, irregularly hexagonal, many somewhat elongate, 10–16 µm, juxtacostal and basal cells of vaginant laminae somewhat larger, quadrate to oblong. |
Seta | to 12 mm. |
4–4.5 mm. |
Sexual condition | rhizoautoicous; perigonia gemmiform, proximal to infertile and fertile stems. |
rhizautoicous or synoicous; perigonia and perichaetia on elongate stems. |
Capsule | theca exserted, ± erect, radially symmetric to inclined, arcuate, bilaterally symmetic, 0.35–1.25 mm; peristome bryoides-type; operculum 0.25–0.3 µm. |
theca exserted, erect, radially symmetric to slightly inclined, bilaterally symmetric, 0.6–0.85 mm; peristome bryoides-type; operculum 0.4 mm. |
Calyptra | cucullate, smooth, 0.5 mm. |
not seen. |
Spores | 11–18 µm. |
14–25 µm. |
Sporophytes | 1 per perichaetium. |
1–2 per perichaetium. |
Fissidens curvatus |
Fissidens appalachensis |
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Habitat | Bare, shaded soil, often among grasses in open Quercus forests and around bases of coastal shrubs | Crevices and surfaces of rocks submerged in swiftly flowing but usually shallow water, sometimes partially emergent |
Distribution |
CA; Mexico; South America; West Indies; Europe; Asia; Africa; Pacific Islands (New Caledonia, New Zealand); Australia |
NC; PA; SC; TN |
Discussion | Fissidens curvatus is distinguished by its delicate dimorphic stems, usually long excurrent costa, limbidium confluent at leaf apex, and slightly elongate laminal cells. The theca is usually arcuate but can also be erect. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens appalachensis, restricted to rapidly moving water, is distinguished by its habitat and strong limbidium. It is most likely to be confused with an expression of F. bryoides that is usually found on wet rocks and stones along the edges of streams, but which differs in its smaller size and weaker limbidium that at times can be partially or completely absent. Laminal cells in size and shape, vaginant laminae, sexuality, thecae, and peristomes in both taxa are similar. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 347. | FNA vol. 27, p. 344. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. milobakeri | |
Name authority | Hornschuch: Linnaea 15: 148. (1841) | R. H. Zander: Bryologist 72: 406, figs. 1–8. (1969) |
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