Fissidens crispus |
Fissidens hyalinus |
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fissidens moss |
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Habit | Plants to 10 × 3 mm. | Plants to 3.5 × 3 mm. |
Stem | unbranched and branched; axillary hyaline nodules absent; central strand present or absent. |
unbranched; axillary hyaline nodules absent; central strand absent. |
Leaves | as many as 20 pairs, elliptic to broadly lanceolate to oblong-lingulate, acute, short-acuminate to obtuse-apiculate, to 2 × 0.5 mm; dorsal lamina narrowed proximally, ending before insertion to ± decurrent; vaginant laminae 2/3–3/4 leaf length, equal; margin ± entire but usually serrulate distally, limbate on all laminae, limbidium usually ending a few cells before apex, frequently edged by 1–2 rows of quadrate to oblong chlorophyllose cells in proximal parts of vaginant laminae, limbidial cells 1–2-stratose; costa percurrent to ending 2–5 cells before leaf, infrequently short-excurrent, bryoides-type; laminal cells 1-stratose, smooth, strongly bulging, densely chlorophyllose, ± obscure, firm-walled, irregularly quadrate to hexagonal, usually arranged in discernable rows in distal part of leaf, 6–10 µm, twice as deep as wide. |
as many as 6 pairs, lanceolate, acute to obtuse, sometimes apiculate, to 2.2 × 0.5 mm; dorsal lamina narrowed proximally, ending at insertion, not decurrent; vaginant laminae 1/3–1/2 leaf length, equal; margin entire but sometimes ± serrulate distally, limbate, limbidium 1–2 cells wide, ending before apex, often indistinct on vaginant laminae, limbidial cells 1-stratose; ecostate, or costa consisting of very short proximal vestige; laminal cells 1-stratose, smooth, thin-walled, hexagonal to elongate-hexagonal, 31–67 × 14–34 µm, ± oblong in proximal portions of leaves, particularly in vaginant laminae. |
Seta | to 9 mm. |
to 3 mm. |
Sexual condition | gonioautoicous and rhizautoicous. |
rhizautoicous; perigonia gemmiform, proximal to perichaetial stems, and on elongate stems, to 1.2 mm. |
Capsule | theca exserted, inclined, bilaterally symmetric to erect, radially symmetric, to 1 mm; peristome bryoides-type; operculum 0.5 mm. |
theca exserted, erect, radially symmetric, sometimes slightly arcuate, bilaterally symmetric, to 0.4 mm; peristome scariosus-type; operculum ± as long as theca. |
Calyptra | cucullate, smooth, 0.8 mm. |
mitrate, prorate, 0.4 mm. |
Spores | 10–16 µm. |
9–13 µm. |
Sporophytes | 1–2 per perichaetium. |
1 per perichaetium. |
Fissidens crispus |
Fissidens hyalinus |
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Habitat | Moist shaded soil, over rocks near streams, infrequently submerged by fluctuating water levels | Bare, clayey soil in shaded ravines, gorges, and dry stream bottoms |
Distribution |
AZ; CA; CO; LA; MS; NM; NV; OR; TX; UT; WA; AB; BC; Mexico; Central America; South America; West Indies |
KY; LA; OH; PA; TN; Asia |
Discussion | Three species belong to the Fissidens crispus complex; F. crispus, F. minutulus, and F. sublimbatus. A well-developed limbidium and small (6–10 µm), bulging, obscure laminal cells that in transverse section are twice as deep as wide characterize all three. Fissidens crispus, better known in western North America as F. limbatus, is highly variable and widespread, ranging widely in tropical America, where it also attains its greatest variability. It is best distinguished by laminal cells that are usually arranged in discernable rows in the distal parts of leaves. The dorsal lamina is quite variable, ending above the insertion to long-decurrent as in some tropical expressions. Leaves, when dry, are usually crispate. The limbidium is found on all or most leaves and usually extends to or ends just before the apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens hyalinus (Latin hyalus, literally “of glass,” alluding to the transparent, glasslike appearance) is doubtless more common than previously thought. B. E. Lemmon (1966) published a photograph showing a perigonial stem and a perichaetial stem with attached sporophyte. A. C. Risk (2002) gave a thorough account of the distribution and habitat of this tiny moss. Plants cannot be mistaken for any other species in North America, owing to the ecostate leaves and smooth, large, thin-walled laminal cells that shrink considerably when dry. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 349. | FNA vol. 27, p. 357. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. limbatus, F. limbatus var. brevifolius, F. limbatus var. ensiformis, F. pusillus var. brevifolius, F. repandus, F. tortilis | |
Name authority | Montagne: Ann. Sci. Nat., Bot., sér. 2, 9: 57. (1838) | Wilson & Hooker f.: J. Bot. (Hooker) 3: 89, fig. 2. (1841) |
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