Fissidens crispus |
Fissidens polypodioides |
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polypody fissidens moss |
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Habit | Plants to 10 × 3 mm. | Plants to 50 × 6.5 mm. |
Stem | unbranched and branched; axillary hyaline nodules absent; central strand present or absent. |
unbranched and sparingly branched; axillary hyaline nodules absent; central strand present. |
Leaves | as many as 20 pairs, elliptic to broadly lanceolate to oblong-lingulate, acute, short-acuminate to obtuse-apiculate, to 2 × 0.5 mm; dorsal lamina narrowed proximally, ending before insertion to ± decurrent; vaginant laminae 2/3–3/4 leaf length, equal; margin ± entire but usually serrulate distally, limbate on all laminae, limbidium usually ending a few cells before apex, frequently edged by 1–2 rows of quadrate to oblong chlorophyllose cells in proximal parts of vaginant laminae, limbidial cells 1–2-stratose; costa percurrent to ending 2–5 cells before leaf, infrequently short-excurrent, bryoides-type; laminal cells 1-stratose, smooth, strongly bulging, densely chlorophyllose, ± obscure, firm-walled, irregularly quadrate to hexagonal, usually arranged in discernable rows in distal part of leaf, 6–10 µm, twice as deep as wide. |
as many as 80 pairs, somewhat undulate, oblong, obtuse to broadly acute, usually bluntly mucronate, to 4 × 1 mm; dorsal lamina rounded proximally, ending at insertion; vaginant laminae 1/2 the leaf length, acute, ± equal, minor lamina ending on or near margin; margin entire but denticulate distally, elimbate; costa percurrent to ending 2–4 cells before apex, taxifolius-type; laminal cells 1-stratose, distinct, smooth, slightly bulging, firm-walled, irregularly hexagonal, 10–20 µm, larger juxtacostally, smaller at margin. |
Seta | to 9 mm. |
to 15 mm. |
Sexual condition | gonioautoicous and rhizautoicous. |
dioicous; perigonia and perichaetia on short axillary branches and main stems. |
Capsule | theca exserted, inclined, bilaterally symmetric to erect, radially symmetric, to 1 mm; peristome bryoides-type; operculum 0.5 mm. |
theca slightly inclined, radially symmetric or slightly arcuate, bilaterally symmetric, to 2.5 mm; operculum about as long as theca. |
Calyptra | cucullate, smooth, 0.8 mm. |
cucullate, smooth, 1.5–1.55 mm. |
Spores | 10–16 µm. |
9–11 µm. |
Sporophytes | 1–2 per perichaetium. |
1–2 per perichaetium. |
Fissidens crispus |
Fissidens polypodioides |
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Habitat | Moist shaded soil, over rocks near streams, infrequently submerged by fluctuating water levels | Calcareous soil, and limestone rocks and boulders along streams and in ravines |
Distribution |
AZ; CA; CO; LA; MS; NM; NV; OR; TX; UT; WA; AB; BC; Mexico; Central America; South America; West Indies |
AL; FL; GA; IN; LA; MS; NC; SC; TX; Mexico; Central America; South America; West Indies; Asia |
Discussion | Three species belong to the Fissidens crispus complex; F. crispus, F. minutulus, and F. sublimbatus. A well-developed limbidium and small (6–10 µm), bulging, obscure laminal cells that in transverse section are twice as deep as wide characterize all three. Fissidens crispus, better known in western North America as F. limbatus, is highly variable and widespread, ranging widely in tropical America, where it also attains its greatest variability. It is best distinguished by laminal cells that are usually arranged in discernable rows in the distal parts of leaves. The dorsal lamina is quite variable, ending above the insertion to long-decurrent as in some tropical expressions. Leaves, when dry, are usually crispate. The limbidium is found on all or most leaves and usually extends to or ends just before the apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens polypodioides is a robust species and has its greatest distribution in the neotropics and Asia. It is perhaps the handsomest species of the genus in the flora area. However, it might be confused with F. asplenioides, from which it is distinguished by taxifolius-type costa, leaves crisped but not curled tightly inward when dry, entire leaf margin, acute and more or less equal vaginant laminae, larger laminal cells that are not lenticularly thickened, and absent differentiated medial marginal cells in the vaginant laminae. The central strand is especially well developed. H. A. Crum and L. E. Anderson (1965, 1981) published illustrations of F. polypodioides and F. asplenioides that aid greatly in distinguishing the two. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 349. | FNA vol. 27, p. 339. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. limbatus, F. limbatus var. brevifolius, F. limbatus var. ensiformis, F. pusillus var. brevifolius, F. repandus, F. tortilis | |
Name authority | Montagne: Ann. Sci. Nat., Bot., sér. 2, 9: 57. (1838) | Hedwig: Sp. Musc. Frond., 154. (1801) |
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