Fissidens asplenioides |
Fissidens subbasilaris |
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asplenium fissidens moss |
fissidens moss |
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Habit | Plants to 250 × 4 mm. | Plants to 12 × 1.5–3.5 mm. |
Stem | unbranched and sparingly branched; axillary hyaline nodules weak; central strand present. |
branched; axillary hyaline nodules absent; central strand present. |
Leaves | in as many as 25 pairs, often undulate, mostly lingulate, rounded to obtuse to broadly acute, sometimes apiculate, to 4 × 0.5 mm; dorsal lamina narrowed proximally, ending before or at insertion, not decurrent; vaginant laminae 1/2–3/4 the leaf length, unequal, minor lamina of most leaves rounded and free distally, or narrowed distally and ending on or near costa; margin ± entire to crenulate-serrulate, sometimes unevenly so distally, elimbate except for a weak limbidium in the proximal parts of vaginant laminae, limbidial cells 1-stratose; costa ending several cells before apex, oblongifolius-type, distal part of leaf in transverse section showing enlarged cells arranged in a single row; laminal cells of dorsal and ventral laminae 1-stratose, distinct, smooth, lenticularly thickened but appearing bulging, firm-walled, irregularly hexagonal, 7–12 µm long; juxtacostal and interior proximal cells of vaginant laminae 1-stratose, smooth, plane, quadrate to ± oblong, larger; medial marginal cells of vaginant laminae ± elongate, oriented obliquely. |
as many as 28 pairs, sometimes ± undulate, oblong, obtuse-apiculate to acute, to 2.4 × 0.3–0.4 mm; dorsal lamina narrowed proximally, ending before or at insertion; vaginant laminae 1/2–2/3 leaf length, acute, ± unequal, minor lamina ending near margin; margin evenly crenulate-serrulate but irregularly serrate distally; costa ending 6–16 cells before apex, covered and obscured above by chlorophyllose cells, taxifolius-type; lamina cells irregularly 2-stratose, smooth, bulging, firm-walled, irregularly rounded-hexagonal, 7.5–10 µm. Sexual condition gonioautoicous and cladautoicous; perigonia gemmiform, axillary, and on short branches; perichaetia on short axillary branches near proximal ends of stem. |
Seta | to 6 mm. |
1.5–5.5 mm. |
Sexual condition | dioicous; perigonia and perichaetia on elongate stems; naked archegonia at times in axils of distal leaves. |
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Capsule | theca ± erect, radially symmetric to slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum ± as long as theca. |
theca 0.8–2.1 mm, erect, radially symmetric or nearly so; peristome taxifolius-type; operculum long rostrate, 0.6–0.9 mm. |
Calyptra | cucullate, smooth, 1–1.7 µm. Spores 7.5–12 µm. |
cucullate, smooth, 1.5 mm. |
Spores | 13–18 µm. |
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Sporophytes | 1–2 per perichaetium, orange-red. |
1 per perichaetium. |
Fissidens asplenioides |
Fissidens subbasilaris |
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Habitat | Sandstone ledges and crevices in moist ravines and grottoes, usually along streams and waterfalls | Particularly around bases of trees, also rocks, mostly limestone, infrequently decaying wood |
Distribution |
AL; GA; KY; LA; MS; NC; SC; Mexico; Central America; South America; West Indies; Asia; Africa; Atlantic Islands (Macaronesia); Pacific Islands (New Zealand); Australia |
AL; AR; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; QC; Mexico (San Luis Potosí, Tamaulipas)
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Discussion | Fissidens asplenioides, usually a robust species, is recognized by leaves typically curled tightly inward from the tips when dry, a minor lamina that in most leaves is rounded distally and attached more or less only along costa, lenticularly thickened dorsal and ventral laminal cells, elongate medial marginal cells of the vaginant laminae oriented obliquely, and oblongifolius-type costa which in the distal part of the leaf in transverse section shows a single row of enlarged cells. The oblongifolius-type costa, found in F. asplenioides and F. santa-clarensis, is unique to sect. Amblyothallia of subg. Pachyfissidens (R. A. Pursell and M. A. Bruggeman-Nannenga 2004). Subterranean, multicellular, irregularly globose, rhizoidal gemmae have been reported in Macaronesian specimens of F. asplenioides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens subbasilaris is distinguished, even in the field, by a costa obscured by chlorophyllose cells distally, and short perichaetial branches in the axils of proximal leaves. The plants are typically dark-green and often tinged with red. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 337. | FNA vol. 27, p. 342. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Name authority | Hedwig: Sp. Musc. Frond., 156. (1801) | Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 6–9. (1801) |
Web links |