Fissidens asplenioides |
Fissidens hallianus |
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asplenium fissidens moss |
fissidens moss |
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Habit | Plants to 250 × 4 mm. | Plants to 120 × 6 mm. |
Stem | unbranched and sparingly branched; axillary hyaline nodules weak; central strand present. |
usually profusely branched; axillary hyaline nodules absent; central strand absent. |
Leaves | in as many as 25 pairs, often undulate, mostly lingulate, rounded to obtuse to broadly acute, sometimes apiculate, to 4 × 0.5 mm; dorsal lamina narrowed proximally, ending before or at insertion, not decurrent; vaginant laminae 1/2–3/4 the leaf length, unequal, minor lamina of most leaves rounded and free distally, or narrowed distally and ending on or near costa; margin ± entire to crenulate-serrulate, sometimes unevenly so distally, elimbate except for a weak limbidium in the proximal parts of vaginant laminae, limbidial cells 1-stratose; costa ending several cells before apex, oblongifolius-type, distal part of leaf in transverse section showing enlarged cells arranged in a single row; laminal cells of dorsal and ventral laminae 1-stratose, distinct, smooth, lenticularly thickened but appearing bulging, firm-walled, irregularly hexagonal, 7–12 µm long; juxtacostal and interior proximal cells of vaginant laminae 1-stratose, smooth, plane, quadrate to ± oblong, larger; medial marginal cells of vaginant laminae ± elongate, oriented obliquely. |
in numerous pairs, lanceolate to linear-lanceolate, acute, to 6.5 × 0.7 mm; dorsal lamina narrowed proximally, usually ending before insertion; vaginant laminae 1/4–1/2 leaf length, acute, unequal in most leaves, minor lamina rounded above, attached on or near costa, ± equal on proximal part of stem, minor lamina ending near leaf margin; margin ± entire, elimbate or limbate in the proximal 2/3 or less of vaginant laminae; costa ending 5–15 cells before apex, bryoides-type; laminal cells 1-stratose, distinct, smooth, slightly bulging, ± firm-walled, quadrate to oblong to hexagonal, 10–15 × 9–11 µm. Sexual condition cladautoicous; perigonia on elongate, axillary branches; perichaetia terminal on mains stems and elongate, axillary branches. |
Seta | to 6 mm. |
0.7–1.5 mm, often bent at the vaginula. |
Sexual condition | dioicous; perigonia and perichaetia on elongate stems; naked archegonia at times in axils of distal leaves. |
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Capsule | theca ± erect, radially symmetric to slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum ± as long as theca. |
theca erect, radially symmetric, 0.4–0.8 mm; peristome anomalous, teeth variable, undivided and often fenestrate to divided 2/3 their length, even in same capsule, nearly smooth to papillose proximally, filaments with straight or spiral thickenings or striations; operculum 0.4–0.6 mm. |
Calyptra | cucullate, smooth, 1–1.7 µm. Spores 7.5–12 µm. |
not seen. |
Spores | 14–32 µm. |
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Sporophytes | 1–2 per perichaetium, orange-red. |
1–2 per perichaetium. |
Fissidens asplenioides |
Fissidens hallianus |
|
Habitat | Sandstone ledges and crevices in moist ravines and grottoes, usually along streams and waterfalls | Flowing and non-flowing water over limestone, granite, and bases of trees, especially abundant in Taxodium-Nyssa swamps |
Distribution |
AL; GA; KY; LA; MS; NC; SC; Mexico; Central America; South America; West Indies; Asia; Africa; Atlantic Islands (Macaronesia); Pacific Islands (New Zealand); Australia |
FL; IL; LA; MA; MS; NC; NJ; TX |
Discussion | Fissidens asplenioides, usually a robust species, is recognized by leaves typically curled tightly inward from the tips when dry, a minor lamina that in most leaves is rounded distally and attached more or less only along costa, lenticularly thickened dorsal and ventral laminal cells, elongate medial marginal cells of the vaginant laminae oriented obliquely, and oblongifolius-type costa which in the distal part of the leaf in transverse section shows a single row of enlarged cells. The oblongifolius-type costa, found in F. asplenioides and F. santa-clarensis, is unique to sect. Amblyothallia of subg. Pachyfissidens (R. A. Pursell and M. A. Bruggeman-Nannenga 2004). Subterranean, multicellular, irregularly globose, rhizoidal gemmae have been reported in Macaronesian specimens of F. asplenioides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens hallianus, named for Elihu Hall, has been confused with F. fontanus; they can be found growing together and both have similar vaginant laminae, although in F. hallianus there can be a weak limbidiuim on the proximal margin. Moreover, the costa is longer in F. hallianus than in F. fontanus. The sporophytes of F. hallianus are terminal, the number per perichaetium usually fewer than in F. fontanus, the seta is longer, relative to the length of the capsule, and the peristome teeth are complete. Sexual reproduction appears to be dependent on seasonal exposure. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 337. | FNA vol. 27, p. 344. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | Conomitrium hallianum, F. manateensis, Octodiceras hallianum | |
Name authority | Hedwig: Sp. Musc. Frond., 156. (1801) | (Sullivant & Lesquereux) Mitten: J. Linn. Soc., Bot. 21: 560. (1885) |
Web links |