Fissidens asplenioides |
Fissidens elegans |
|
---|---|---|
asplenium fissidens moss |
elegant fissidens moss |
|
Habit | Plants to 250 × 4 mm. | Plants to 5.5 × 2 mm. |
Stem | unbranched and sparingly branched; axillary hyaline nodules weak; central strand present. |
unbranched and branched; axillary hyaline nodules absent; central strand weak. |
Leaves | in as many as 25 pairs, often undulate, mostly lingulate, rounded to obtuse to broadly acute, sometimes apiculate, to 4 × 0.5 mm; dorsal lamina narrowed proximally, ending before or at insertion, not decurrent; vaginant laminae 1/2–3/4 the leaf length, unequal, minor lamina of most leaves rounded and free distally, or narrowed distally and ending on or near costa; margin ± entire to crenulate-serrulate, sometimes unevenly so distally, elimbate except for a weak limbidium in the proximal parts of vaginant laminae, limbidial cells 1-stratose; costa ending several cells before apex, oblongifolius-type, distal part of leaf in transverse section showing enlarged cells arranged in a single row; laminal cells of dorsal and ventral laminae 1-stratose, distinct, smooth, lenticularly thickened but appearing bulging, firm-walled, irregularly hexagonal, 7–12 µm long; juxtacostal and interior proximal cells of vaginant laminae 1-stratose, smooth, plane, quadrate to ± oblong, larger; medial marginal cells of vaginant laminae ± elongate, oriented obliquely. |
as many as 12 pairs, oblong to lanceolate, acute to obtuse-apiculate, most ending in a clear, sharp cell, to 1.8 × 0.4 mm; dorsal lamina narrowed or rounded proximally, ending at or before insertion, not decurrent; vaginant laminae 1/2–2/3 leaf length, unequal, minor lamina ending near margin; margin serrulate, limbate and entire to remotely denticulate on proximal 2/3 or less of vaginant laminae of most leaves of perichaetial stems, often absent from leaves of infertile stems, limbidium intralaminal in part or completely, limbidial cells 1-stratose; costa ending 2–4 cells before apex or percurrent, ending in apiculus, infrequently short-excurrent, bryoides-type; lamina cells 1-stratose, pluripapillose, obscure, firm-walled, rounded-hexagonal, 5–8 µm, twice as deep as wide. |
Seta | to 6 mm. |
to 5 mm. |
Sexual condition | dioicous; perigonia and perichaetia on elongate stems; naked archegonia at times in axils of distal leaves. |
rhizautoicous and gonioautoicous. |
Capsule | theca ± erect, radially symmetric to slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum ± as long as theca. |
theca exserted, erect, radially symmetric, to 0.8 mm; peristome scariosus-type; operculum to 0.5 mm. |
Calyptra | cucullate, smooth, 1–1.7 µm. Spores 7.5–12 µm. |
cucullate, smooth, to 0.6 mm. |
Spores | 9–13 µm. |
|
Sporophytes | 1–2 per perichaetium, orange-red. |
1 per perichaetium. |
Fissidens asplenioides |
Fissidens elegans |
|
Habitat | Sandstone ledges and crevices in moist ravines and grottoes, usually along streams and waterfalls | Sandy and clayey soils along roadsides and streams, banks of ravines, bluffs, loess banks, uprooted tree roots, trunks of trees, decaying stumps, sandstone |
Distribution |
AL; GA; KY; LA; MS; NC; SC; Mexico; Central America; South America; West Indies; Asia; Africa; Atlantic Islands (Macaronesia); Pacific Islands (New Zealand); Australia |
AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; SC; TN; TX; VA; Mexico; Central America; South America; West Indies
|
Discussion | Fissidens asplenioides, usually a robust species, is recognized by leaves typically curled tightly inward from the tips when dry, a minor lamina that in most leaves is rounded distally and attached more or less only along costa, lenticularly thickened dorsal and ventral laminal cells, elongate medial marginal cells of the vaginant laminae oriented obliquely, and oblongifolius-type costa which in the distal part of the leaf in transverse section shows a single row of enlarged cells. The oblongifolius-type costa, found in F. asplenioides and F. santa-clarensis, is unique to sect. Amblyothallia of subg. Pachyfissidens (R. A. Pursell and M. A. Bruggeman-Nannenga 2004). Subterranean, multicellular, irregularly globose, rhizoidal gemmae have been reported in Macaronesian specimens of F. asplenioides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The obscure, pluripapillose lamina cells and leaf apex that terminates in a single, hyaline, sharply pointed cell distinguish Fissidens elegans. The limbidium is quite variable, occurring on the vaginant laminae of most leaves of perichaetial stems to being absent from the leaves of infertile stems. The species is close to F. pallidinervis (see discussion under 26). H. A. Crum and L.E. Anderson (1981) commented on a collection that was possibly made in Lafayette, Wisconsin, a site quite distant from the normal distribution of this species in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 337. | FNA vol. 27, p. 350. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Synonyms | F. ravenelii | |
Name authority | Hedwig: Sp. Musc. Frond., 156. (1801) | Bridel: Muscol. Recent., suppl. 1: 167. (1806) |
Web links |