Fissidens asplenioides |
Fissidens appalachensis |
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asplenium fissidens moss |
Appalachian fissidens moss |
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Habit | Plants to 250 × 4 mm. | Plants to 13 × 2–4 mm. |
Stem | unbranched and sparingly branched; axillary hyaline nodules weak; central strand present. |
unbranched and branched; axillary hyaline nodules weak or absent; central strand weak. |
Leaves | in as many as 25 pairs, often undulate, mostly lingulate, rounded to obtuse to broadly acute, sometimes apiculate, to 4 × 0.5 mm; dorsal lamina narrowed proximally, ending before or at insertion, not decurrent; vaginant laminae 1/2–3/4 the leaf length, unequal, minor lamina of most leaves rounded and free distally, or narrowed distally and ending on or near costa; margin ± entire to crenulate-serrulate, sometimes unevenly so distally, elimbate except for a weak limbidium in the proximal parts of vaginant laminae, limbidial cells 1-stratose; costa ending several cells before apex, oblongifolius-type, distal part of leaf in transverse section showing enlarged cells arranged in a single row; laminal cells of dorsal and ventral laminae 1-stratose, distinct, smooth, lenticularly thickened but appearing bulging, firm-walled, irregularly hexagonal, 7–12 µm long; juxtacostal and interior proximal cells of vaginant laminae 1-stratose, smooth, plane, quadrate to ± oblong, larger; medial marginal cells of vaginant laminae ± elongate, oriented obliquely. |
as many as 22 pairs, ligulate to oblong-lanceolate, obtuse-mucronate to acute, to 4 × 0.3–0.5 mm; dorsal lamina narrowed proximally, ending at insertion or slightly before; vaginant laminae 1/2–2/3 leaf length, equal in proximal leaves, unequal in distal leaves, minor lamina ending between margin and costa or rounded and ending on or near costa; margin entire but sometimes serrulate distally, limbate on all laminae, limbidium confluent at apex or ending a few cells before, limbidial cells 2- to 5-stratose; costa percurrent or ending in mucro, bryoides-type; laminal cells 1- to irregularly 2-stratose, smooth, slightly bulging, firm-walled, irregularly hexagonal, many somewhat elongate, 10–16 µm, juxtacostal and basal cells of vaginant laminae somewhat larger, quadrate to oblong. |
Seta | to 6 mm. |
4–4.5 mm. |
Sexual condition | dioicous; perigonia and perichaetia on elongate stems; naked archegonia at times in axils of distal leaves. |
rhizautoicous or synoicous; perigonia and perichaetia on elongate stems. |
Capsule | theca ± erect, radially symmetric to slightly arcuate, bilaterally symmetric, to 1.5 mm; peristome taxifolius-type; operculum ± as long as theca. |
theca exserted, erect, radially symmetric to slightly inclined, bilaterally symmetric, 0.6–0.85 mm; peristome bryoides-type; operculum 0.4 mm. |
Calyptra | cucullate, smooth, 1–1.7 µm. Spores 7.5–12 µm. |
not seen. |
Spores | 14–25 µm. |
|
Sporophytes | 1–2 per perichaetium, orange-red. |
1–2 per perichaetium. |
Fissidens asplenioides |
Fissidens appalachensis |
|
Habitat | Sandstone ledges and crevices in moist ravines and grottoes, usually along streams and waterfalls | Crevices and surfaces of rocks submerged in swiftly flowing but usually shallow water, sometimes partially emergent |
Distribution |
AL; GA; KY; LA; MS; NC; SC; Mexico; Central America; South America; West Indies; Asia; Africa; Atlantic Islands (Macaronesia); Pacific Islands (New Zealand); Australia |
NC; PA; SC; TN |
Discussion | Fissidens asplenioides, usually a robust species, is recognized by leaves typically curled tightly inward from the tips when dry, a minor lamina that in most leaves is rounded distally and attached more or less only along costa, lenticularly thickened dorsal and ventral laminal cells, elongate medial marginal cells of the vaginant laminae oriented obliquely, and oblongifolius-type costa which in the distal part of the leaf in transverse section shows a single row of enlarged cells. The oblongifolius-type costa, found in F. asplenioides and F. santa-clarensis, is unique to sect. Amblyothallia of subg. Pachyfissidens (R. A. Pursell and M. A. Bruggeman-Nannenga 2004). Subterranean, multicellular, irregularly globose, rhizoidal gemmae have been reported in Macaronesian specimens of F. asplenioides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Fissidens appalachensis, restricted to rapidly moving water, is distinguished by its habitat and strong limbidium. It is most likely to be confused with an expression of F. bryoides that is usually found on wet rocks and stones along the edges of streams, but which differs in its smaller size and weaker limbidium that at times can be partially or completely absent. Laminal cells in size and shape, vaginant laminae, sexuality, thecae, and peristomes in both taxa are similar. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 337. | FNA vol. 27, p. 344. |
Parent taxa | Fissidentaceae > Fissidens | Fissidentaceae > Fissidens |
Sibling taxa | ||
Name authority | Hedwig: Sp. Musc. Frond., 156. (1801) | R. H. Zander: Bryologist 72: 406, figs. 1–8. (1969) |
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