Festuca rubra |
Festuca viviparoidea |
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fetuque rouge, ravine fescue, red fescue |
northern fescue, viviparous fescue |
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Habit | Plants usually rhizomatous, usually loosely to densely cespitose, culms sometimes single and widely spaced, sometimes stoloniferous. | Plants loosely or densely cespitose, without rhizomes. | ||||||||||||||||||||||||||||||||||||||||||||||||
Culms | (8)10-120 (130) cm, erect or decumbent, glabrous and smooth. |
(11)13.5-25(28) cm, smooth and glabrous throughout or sparsely to densely scabrous or puberulent below the inflorescence. |
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Sheaths | closed for about 3/4 their length when young, readily splitting with age, usually pubescent, at least distally, hairs retrorse or antrorse, sometimes glabrous, not persistent, older vegetative shoot sheaths shredding into fibers; collars glabrous; ligules 0.1-0.5 mm; blades usually conduplicate or convolute and 0.3-2.5 mm in diameter, sometimes flat and 1.5-7 mm wide, abaxial surfaces glabrous, smooth or scabrous, adaxial surfaces scabrous or pubescent, veins 5-9(13), ribs (3)5-7(9), usually conspicuous; abaxial sclerenchyma in 5-9(13) discrete or partly confluent strands, rarely forming a complete band; adaxial sclerenchyma sometimes present in fascicles opposite the veins; girders and pillars not developed. |
closed for about 1/2 their length, glabrous or scabrous, stramineous or brownish, persistent or slowly shredding into fibers; collars glabrous; ligules 0.1-0.5 mm; blades 0.5-1 mm in diameter, conduplicate, abaxial surfaces glabrous, smooth or scabrous, adaxial surfaces scabrous, veins 5-7, ribs 3-5, 1 distinct and 2-4 indistinct; abaxial sclerenchyma in 3-7 small strands, covering less than 1/2 the abaxial surface and usually less than twice as wide as high. |
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Inflorescences | (2)3.5-25(30) cm, usually open or loosely contracted panicles, occasionally racemes, with 1-3 branches per node, lower branches with 2+ spikelets; branches erect or spreading, stiff or lax, glabrous, scabrous, or pubescent. |
(1)3-4.8 cm, contracted, usually panicles, sometimes racemes, erect, with 1-2 branches per node; branches erect, lower branches with (1)2+ spikelets. |
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Spikelets | (6)7-17 mm, with 3-10 florets. |
pseudoviviparous, their length varying with the stage of vegetative proliferation, the glumes and often 1 or 2 adjacent florets more or less normally developed, or only slightly elongated, the distal florets replaced by bracts. |
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Glumes | ovate-lanceolate to lanceolate, exceeded by the distal florets; lower glumes (1.5)2-6(7) mm; upper glumes (3)3.5-8.5 mm; lemmas 4-9.5 mm, usually glabrous and smooth, sometimes scabrous towards the apices, sometimes densely pubescent throughout, attenuate or acuminate in side view, awned, awns (0.1)0.4-4.5 mm; paleas slightly shorter than to about equaling the lemmas, intercostal region puberulent distally; anthers 1.8-4.5 mm; ovary apices glabrous. |
lanceolate, glabrous and smooth, sometimes scabrous towards the apices, or puberulent throughout or only towards the apices; lower glumes (2)3-6 mm; upper glumes (2.7)3-7 mm; normal lemmas 3.3-6 mm, mostly smooth or scabrous distally, glabrous or puberulent, awned or unawned, sometimes varying within a panicle, awns to 1 mm; vegetative bracts unawned, leaflike, sometimes with ligules; paleas usually reduced or absent, well-formed paleas about as long as the lemmas, intercostal region scabrous or puberulent distally; anthers usually not developed, well-formed anthers to about 2 mm; ovaries sometimes not developed; ovary apices, when present, glabrous. |
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2n | = 28, 42, 56, 70. |
= 49, 56. |
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Festuca rubra |
Festuca viviparoidea |
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Distribution |
AK; AL; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
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AK; MT; WY; AB; BC; NL; NT; NU; QC; YT; Greenland |
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Discussion | Festuca rubra is interpreted here as a morphologically diverse polyploid complex that is widely distributed in the arctic and temperate zones of Europe, Asia, and North America. Its treatment is complicated by the fact that Eurasian material has been introduced in other parts of the world. In addition, hundreds of forage and turf cultivars have been developed, many of which have also been widely distributed. Within the complex, morphologically, ecologically, geographically, and/or cytologically distinct taxa have been described, named, and given various taxonomic ranks. In some cases, these taxa represent extremes, and in other cases they are morphologically intermediate between other taxa. Moreover, hybridization and/or introgression between native taxa, and between native and non-native taxa, may be occurring. In Iceland and southern Greenland, putative hybrids between Festuca frederikseniae and F. rubra have been reported, and named F. villosa-vivipara (Rosenv.) E.B. Alexeev (see under F. frederikseniae, p. 436). Overlap in morphological characters between most taxa in the complex has led some taxonomists to ignore the variation within the complex, calling all its members Festuca rubra without qualification. This obscures what is known about the complex, and presents an extremely heterogenous assemblage of plants as a single “species”—or a mega-species. The following account attempts to reflect the genetic diversity of the F. rubra complex in the Flora region. All the taxa are recognized as subspecies, but they are not necessarily equivalent in terms of their distinction and genetic isolation. Much more work on the taxonomy of the F. rubra complex is needed before the boundairs of individual taxa can be firmly established. Some variants that need attention are (1) plants growing on the sandy shores of the Great Lakes that have glaucous leaves and spikelets, sometimes treated as F. rubra var. juncea (Hack.) K. Richt., (2) native plants along the James Bay and Hudson Bay shore that are ecologically distinct from F. rubra subsp. rubra, (3) native plants growing in marshes, sometimes called F. rubra var. megastachys (Gaudin) Hegi (Dore and McNeill 1980), (4) seashore variants along the Atlantic coast of North America, (5) plants with glaucescent leaves and spikelets which are widely distributed in the Flora region and have been called F. rubra subsp. glaucodea Piper, (6) the widespread variant with pubescent to villous lemmas, sometimes called F. rubra f. squarrosa (Hartm.) Holmb. Festuca earlei (p. 420) is sometimes confused with F. rubra. It differs in having pubescent ovary apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Festuca viviparoidea is circumboreal in distribution. It may consist of hybrids between Festuca baffinensis (p. 432) and F. brachyphylla (p. 428) and/or other species (see under F. frederikseniae, above). It has frequently been included in F. ovina (p. 422). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 412. | FNA vol. 24, p. 436. | ||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | F. vivipara subsp. glabra | |||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | L. | Krajina ex Pavlick | ||||||||||||||||||||||||||||||||||||||||||||||||
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