Festuca rubra |
Festuca minutiflora |
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fetuque rouge, ravine fescue, red fescue |
little fescue, small-flower fescue |
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Habit | Plants usually rhizomatous, usually loosely to densely cespitose, culms sometimes single and widely spaced, sometimes stoloniferous. | Plants loosely or densely cespitose, without rhizomes. | ||||||||||||||||||||||||||||||||||||||||||||
Culms | (8)10-120 (130) cm, erect or decumbent, glabrous and smooth. |
4-30 cm, usually erect, sometimes semi-prostrate, glabrous, smooth. |
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Sheaths | closed for about 3/4 their length when young, readily splitting with age, usually pubescent, at least distally, hairs retrorse or antrorse, sometimes glabrous, not persistent, older vegetative shoot sheaths shredding into fibers; collars glabrous; ligules 0.1-0.5 mm; blades usually conduplicate or convolute and 0.3-2.5 mm in diameter, sometimes flat and 1.5-7 mm wide, abaxial surfaces glabrous, smooth or scabrous, adaxial surfaces scabrous or pubescent, veins 5-9(13), ribs (3)5-7(9), usually conspicuous; abaxial sclerenchyma in 5-9(13) discrete or partly confluent strands, rarely forming a complete band; adaxial sclerenchyma sometimes present in fascicles opposite the veins; girders and pillars not developed. |
closed for about 1/2 their length, glabrous, persistent; collars glabrous; ligules 0.1-0.3 mm; blades (0.2)0.3-0.4(0.6) mm in diameter, conduplicate, lax, abaxial surfaces glabrous, adaxial surfaces sparsely scabrous to puberulent, veins 3-5, ribs 1-3; abaxial sclerenchyma in 3-5 small strands, less than twice as wide as high; adaxial sclerenchyma absent. |
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Inflorescences | (2)3.5-25(30) cm, usually open or loosely contracted panicles, occasionally racemes, with 1-3 branches per node, lower branches with 2+ spikelets; branches erect or spreading, stiff or lax, glabrous, scabrous, or pubescent. |
1-4(5) cm, contracted, with 1-2 branches per node; branches erect, lower branches with 2+ spikelets. |
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Spikelets | (6)7-17 mm, with 3-10 florets. |
(2.5)3-5 mm, with (1)2-3(5) florets. |
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Glumes | ovate-lanceolate to lanceolate, exceeded by the distal florets; lower glumes (1.5)2-6(7) mm; upper glumes (3)3.5-8.5 mm; lemmas 4-9.5 mm, usually glabrous and smooth, sometimes scabrous towards the apices, sometimes densely pubescent throughout, attenuate or acuminate in side view, awned, awns (0.1)0.4-4.5 mm; paleas slightly shorter than to about equaling the lemmas, intercostal region puberulent distally; anthers 1.8-4.5 mm; ovary apices glabrous. |
exceeded by the upper florets, ovate to ovate-lanceolate, sparsely scabrous distally; lower glumes 1.3-2.5 mm; upper glumes 2-3.5 mm; lemmas (2)2.2-3.5(4) mm, ovate-lanceolate, sparsely scabrous near the apices, apices abruptly acuminate, awns 0.5-1.5(1.7) mm; paleas about as long as or slightly shorter than the lemmas, intercostal region scabrous distally; anthers (0.4)0.6-1.2 mm; ovary apices usually with a few hairs, rarely glabrous. |
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Flag | leaf blades 0.7-3.5 cm. |
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2n | = 28, 42, 56, 70. |
= 28. |
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Festuca rubra |
Festuca minutiflora |
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Distribution |
AK; AL; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; TN; TX; UT; VA; VT; WA; WI; WV; WY; HI; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT; Greenland
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AK; AZ; CA; CO; ID; MT; NM; NV; OR; UT; WY; AB; BC; YT |
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Discussion | Festuca rubra is interpreted here as a morphologically diverse polyploid complex that is widely distributed in the arctic and temperate zones of Europe, Asia, and North America. Its treatment is complicated by the fact that Eurasian material has been introduced in other parts of the world. In addition, hundreds of forage and turf cultivars have been developed, many of which have also been widely distributed. Within the complex, morphologically, ecologically, geographically, and/or cytologically distinct taxa have been described, named, and given various taxonomic ranks. In some cases, these taxa represent extremes, and in other cases they are morphologically intermediate between other taxa. Moreover, hybridization and/or introgression between native taxa, and between native and non-native taxa, may be occurring. In Iceland and southern Greenland, putative hybrids between Festuca frederikseniae and F. rubra have been reported, and named F. villosa-vivipara (Rosenv.) E.B. Alexeev (see under F. frederikseniae, p. 436). Overlap in morphological characters between most taxa in the complex has led some taxonomists to ignore the variation within the complex, calling all its members Festuca rubra without qualification. This obscures what is known about the complex, and presents an extremely heterogenous assemblage of plants as a single “species”—or a mega-species. The following account attempts to reflect the genetic diversity of the F. rubra complex in the Flora region. All the taxa are recognized as subspecies, but they are not necessarily equivalent in terms of their distinction and genetic isolation. Much more work on the taxonomy of the F. rubra complex is needed before the boundairs of individual taxa can be firmly established. Some variants that need attention are (1) plants growing on the sandy shores of the Great Lakes that have glaucous leaves and spikelets, sometimes treated as F. rubra var. juncea (Hack.) K. Richt., (2) native plants along the James Bay and Hudson Bay shore that are ecologically distinct from F. rubra subsp. rubra, (3) native plants growing in marshes, sometimes called F. rubra var. megastachys (Gaudin) Hegi (Dore and McNeill 1980), (4) seashore variants along the Atlantic coast of North America, (5) plants with glaucescent leaves and spikelets which are widely distributed in the Flora region and have been called F. rubra subsp. glaucodea Piper, (6) the widespread variant with pubescent to villous lemmas, sometimes called F. rubra f. squarrosa (Hartm.) Holmb. Festuca earlei (p. 420) is sometimes confused with F. rubra. It differs in having pubescent ovary apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Festuca minutiflora grows in alpine regions of the western mountains, from southeastern Alaska and the southwestern Yukon Territory to Arizona, New Mexico, and the Sierra Nevada of California. It has often been overlooked or included with F. brachyphylla (p. 428), from which it differs in its laxer and narrower leaves, looser panicles, smaller spikelets, more pointed lemmas, shorter awns, and scattered hairs on the ovary. In the southern Rocky Mountains, it may grow with F. earlei (p. 420), which has short rhizomes and larger spikelets and lemmas. Festuca minutiflora has frequently been included in F. ovina (p. 422). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 412. | FNA vol. 24, p. 434. | ||||||||||||||||||||||||||||||||||||||||||||
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Name authority | L. | Rydb. | ||||||||||||||||||||||||||||||||||||||||||||
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