Festuca californica |
Poaceae subfam. pooideae |
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California fescue |
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Habit | Plants densely cespitose, without rhizomes. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||||||||||
Culms | 30-150 (200) cm, glabrous or pubescent, sometimes scabrous. |
usually hollow, sometimes solid. |
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Sheaths | closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins; collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous; ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent; blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17); abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands; adaxial sclerenchyma sometimes present; girders or pillars present at most veins. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Inflorescences | 10-25(30) cm, open, with (1)2(4) branches per node; branches spreading and lax. |
usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | 8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | lanceolate, glabrous or sparsely scabrous at the apices; lower glumes (4)4.5-6.7(8) mm; upper glumes (5)6-10 mm; lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm; paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally; anthers (3)4-7.5(8.5) mm; ovary apices densely pubescent. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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2n | = 56. |
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Festuca californica |
Poaceae subfam. pooideae |
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Distribution |
CA; OR
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Discussion | Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 410. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae | Poaceae | ||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Vasey | Benth. | ||||||||||||||||||||||||||||||||||||||||||||
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