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California fescue

Howell's fescue, Washington fescue

Habit Plants densely cespitose, without rhizomes. Plants loosely or densely cespitose, without rhizomes.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

40-70(100) cm, smooth, glabrous throughout;

nodes usually not exposed.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for less than 1/2 their length, glabrous or scabrous, persistent or slowly shredding into fibers;

collars glabrous;

ligules (0.2)0.3-0.5 mm;

blades 1.5-3 mm in diameter, loosely conduplicate to flat, persistent, abaxial surfaces glabrous and smooth, adaxial surfaces scabrous or pubescent on the ribs, veins 7-13, ribs 7-10(13), blades of the lower and upper cauline leaves similar in length and stiffness;

abaxial sclerenchyma in strands opposite and about as wide as the major veins;

adaxial sclerenchyma often present opposite the major veins;

pillars or girders often developed.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

8-12(15) cm, loosely contracted, with 1-2 branches per node;

branches lax, spreading or loosely erect, lower branches with 2+ spikelets.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

8-15(18) mm, with (3)4-6(10) florets.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

exceeded by the upper florets, lanceolate to ovate-lanceolate, scabrous distally;

lower glumes (2)3.5-5.5 mm;

upper glumes (4)5.5-7(8) mm;

calluses wider than long, glabrous, sometimes slightly scabrous;

lemmas (5.5)8-10(11) mm, lanceolate, scabrous or puberulent at least distally, attenuate, sometimes minutely bidentate, awns 1-3(3.5) mm, terminal or subterminal, straight, occasionally absent;

paleas about as long as the lemmas, intercostal region scabrous or puberulent distally;

anthers (3)3.7-5.7 mm;

ovary apices sparsely or densely pubescent.

2n

= 56.

= unknown.

Festuca californica

Festuca washingtonica

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
WA; BC
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca washingtonica grows in subalpine to low alpine regions of British Columbia and Washington. It has also been reported from Oregon and northern California; these records have not been verified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
Source FNA vol. 24, p. 410. FNA vol. 24, p. 440.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
Name authority Vasey E.B. Alexeev
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