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California fescue

northern fescue, viviparous fescue

Habit Plants densely cespitose, without rhizomes. Plants loosely or densely cespitose, without rhizomes.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

(11)13.5-25(28) cm, smooth and glabrous throughout or sparsely to densely scabrous or puberulent below the inflorescence.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for about 1/2 their length, glabrous or scabrous, stramineous or brownish, persistent or slowly shredding into fibers;

collars glabrous;

ligules 0.1-0.5 mm;

blades 0.5-1 mm in diameter, conduplicate, abaxial surfaces glabrous, smooth or scabrous, adaxial surfaces scabrous, veins 5-7, ribs 3-5, 1 distinct and 2-4 indistinct;

abaxial sclerenchyma in 3-7 small strands, covering less than 1/2 the abaxial surface and usually less than twice as wide as high.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

(1)3-4.8 cm, contracted, usually panicles, sometimes racemes, erect, with 1-2 branches per node;

branches erect, lower branches with (1)2+ spikelets.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

pseudoviviparous, their length varying with the stage of vegetative proliferation, the glumes and often 1 or 2 adjacent florets more or less normally developed, or only slightly elongated, the distal florets replaced by bracts.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

lanceolate, glabrous and smooth, sometimes scabrous towards the apices, or puberulent throughout or only towards the apices;

lower glumes (2)3-6 mm;

upper glumes (2.7)3-7 mm;

normal lemmas 3.3-6 mm, mostly smooth or scabrous distally, glabrous or puberulent, awned or unawned, sometimes varying within a panicle, awns to 1 mm;

vegetative bracts unawned, leaflike, sometimes with ligules;

paleas usually reduced or absent, well-formed paleas about as long as the lemmas, intercostal region scabrous or puberulent distally;

anthers usually not developed, well-formed anthers to about 2 mm;

ovaries sometimes not developed;

ovary apices, when present, glabrous.

2n

= 56.

= 49, 56.

Festuca californica

Festuca viviparoidea

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MT; WY; AB; BC; NL; NT; NU; QC; YT; Greenland
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca viviparoidea is circumboreal in distribution. It may consist of hybrids between Festuca baffinensis (p. 432) and F. brachyphylla (p. 428) and/or other species (see under F. frederikseniae, above). It has frequently been included in F. ovina (p. 422).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
1. Plants loosely cespitose; culms usually glabrous and smooth throughout, rarely sparsely puberulent near the inflorescence; sheaths brownish, slowly shredding into fibers; abaxial sclerenchyma strands less than 2 times as wide as high; glumes and lemmas puberulent throughout or only near the apices
Festuca viviparoidea subsp. krajinae
1. Plants densely cespitose; culms densely to sparsely puberulent below the inflorescence; shearhs stramineous, persistent; abaxial sclerenchyma strands 2-3 times wider than high; glumes and lemmas smooth or scabrous near the apices
subsp. viviparoidea
Source FNA vol. 24, p. 410. FNA vol. 24, p. 436.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
F. viviparoidea subsp. viviparoidea
Synonyms F. vivipara subsp. glabra
Name authority Vasey Krajina ex Pavlick
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