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California fescue

fétuque dressée à feuilles scabres, hard fescue, hard or sheep fescue, rough leaf fescue, sheep fescue

Habit Plants densely cespitose, without rhizomes. Plants densely cespitose, without rhizomes.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

(15)20-75 (80) cm, smooth, glabrous or with sparse hairs.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for less than 1/3 their length, usually glabrous, rarely pubescent, persistent;

collars glabrous;

ligules 0.1-0.5 mm;

blades (0.5)0.8-1.2 mm in diameter, usually conduplicate, rarely flat, abaxial surfaces glabrous, puberulent, or scabrous, adaxial surfaces scabrous or puberulent to pubescent, veins 5-7(9), ribs 3-7, usually distinct;

abaxial sclerenchyma usually in an irregular, interrupted or continuous band, rarely in 5-7 small strands, usually more than twice as wide as high;

adaxial sclerenchyma absent.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

(2.5)3-13(16) cm, contracted, with 1-2 branches per node;

branches erect or stiffly spreading, secondary branches not divaricate, lower branches with 2+ spikelets.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

5-9(10.8) mm, with 3-7(8) florets.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

exceeded by the upper florets, ovate-lanceolate to lanceolate, mostly smooth and glabrous, sometimes scabrous and/or pubescent distally;

lower glumes (1.8)2-3.5(4) mm;

upper glumes 3-5(5.5) mm;

lemmas 3.8-5(6.5) mm, lanceolate, usually smooth and glabrous on the lower portion and scabrous or pubescent distally, especially on the margins, rarely entirely pubescent, awns 0.5-2.5(3) mm, usually less than 1/2 as long as the lemma body;

paleas about as long as the lemmas, intercostal region puberulent distally;

anthers (1.8)2.3-3.4 mm;

ovary apices glabrous.

2n

= 56.

= 42.

Festuca californica

Festuca trachyphylla

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from USDA
[WildflowerSearch map]
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca trachyphylla is native to open forests and forest edge habitats of Europe. It has been introduced and has become naturalized in many temperate regions. In the Flora region, F. trachyphylla is generally sold under the name 'Hard Fescue', and is popular as a durable turf grass and soil stabilizer. It is particularly common in the eastern United States and southeastern Canada, but is probably grown throughout the temperate parts of the region. Its naturalized distribution can be expected to expand.

For many years, Festuca trachyphylla was known, inappropriately, under other names, e.g., F. duriuscula L., F. ovina var. duriuscula (L.) W.D.J. Koch, and F. longifolia Thuill. Some European authors treat it as F. stricta subsp. trachyphylla (Hack.) Patzke. It has frequently been included in F. ovina (p. 422).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
Source FNA vol. 24, p. 410. FNA vol. 24, p. 424.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
Synonyms F. ovina var. duriuscula, F. longifolia var. trachyphylla, F. duriuscula, F. brevipila
Name authority Vasey (Hack.) Krajina
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