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California fescue

ravine fescue

Habit Plants densely cespitose, without rhizomes. Plants loosely cespitose, without rhizomes, with short extra-vaginal tillers.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

60-100 (150) cm, erect, glabrous;

nodes usually exposed.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for less than 1/3 their length, glabrous or scabrous, shredding into fibers;

ligules 0.3-1.5 mm;

blades 3-6(10) mm wide, flat, lax, margins scabrous, abaxial and adaxial surfaces glabrous, adaxial surfaces sometimes scabrous, veins 13-25, ribs obscure;

abaxial sclerenchyma in narrow strands;

adaxial sclerenchyma developed;

girders or pillars formed at the major veins.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

10-20(40) cm, open or somewhat contracted, with 1-2(3) branches per node;

branches lax, more or less spreading, spikelets borne towards the ends of the branches.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

7-12 mm, with (2)3-5 florets.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

lanceolate, scabrous at least on the midvein, acute to acuminate;

lower glumes (1.5) 2.5-4(4.5) mm;

upper glumes (3)4-6.5 mm;

calluses wider than long, glabrous, smooth or slightly scabrous;

lemmas (5)6-8(9) mm, lanceolate, scabrous or puberulent, acuminate, unawned or awned, awns to 2 mm;

paleas about as long as or shorter than the lemmas, intercostal region puberulent distally;

anthers 1.6-2.5 mm;

ovary apices pubescent.

2n

= 56.

= unknown.

Festuca californica

Festuca sororia

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; MO; NM; UT
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca sororia grows in open woods and on shaded slopes and stream banks, at 2000-3000 m. It is restricted to the United States, growing from central Utah and Colorado to Arizona and New Mexico. A single puzzling specimen is the basis for the reported occurrence of this species in Missouri (Yatskievych 1999).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
Source FNA vol. 24, p. 410. FNA vol. 24, p. 404.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Subulatae > sect. Subulatae
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
Name authority Vasey Piper
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