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California fescue

fescue, pseudoviviparous fescue, pseusoviviparous fescue

Habit Plants densely cespitose, without rhizomes. Plants loosely cespitose, rhizomatous.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

30-60 cm.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for about 3/4 their length, glabrous or scabrous-pubescent, shredding into fibers, bases red-scarious;

collars glabrous;

ligules 0.5-1 mm long;

vegetative shoot blades to about 2 mm wide when flat, 0.5-1 mm in diameter when loosely conduplicate, deep green, abaxial surfaces more or less uniformly scabrous, adaxial surfaces hispid or pilose on the ribs;

abaxial sclerenchyma in 5-9 small strands;

adaxial sclerenchyma absent;

cauline blades 1.4-2.5 mm wide, flat.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

(4)7-12(15) cm, open, lax, secund or partially secund, with 1-2 branches per node;

branches somewhat stiff or lax, lower branches with 2-5 spikelets.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

pseudoviviparous, varying in length with the stage of vegetative proliferation, most florets replaced by bracts, the glumes and sometimes the lowest floret more or less normally developed or only slightly elongated, mostly deep green or reddish tinged.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

more or less normally developed, lanceolate, apices scabrous;

lower glumes (2.5)3.5-6 mm;

upper glumes 4.5-6.5(8) mm;

lemmas and bracts glabrous or pubescent, smooth or scabrous, sometimes mucronate, mucros to 0.5 mm;

paleas, if present, about as long as or shorter than the lemmas, intercostal region puberulent distally;

anthers not developed or abortive, to 2 mm;

ovaries not developed.

2n

= 56.

= ca. 70.

Festuca californica

Festuca pseudovivipara

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
BC
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca pseudovivipara grows on coastal mountainsides, scree slopes, and other rocky areas, at 300-800 m. It is known only from the Queen Charlotte Islands, British Columbia.

Festuca pseudovivipara has been described as a form of F. rubra subsp. aucta (p. 414), but differs from that taxon in having pseudoviviparous spikelets. It is also ecologically, altitudinally, and probably reproductively isolated from F. rubra subsp. aucta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
Source FNA vol. 24, p. 410. FNA vol. 24, p. 419.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
Name authority Vasey (Pavlick) Pavlick
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