Festuca californica |
Festuca prolifera |
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California fescue |
proliferous fescue |
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Habit | Plants densely cespitose, without rhizomes. | Plants usually loosely cespitose, often mat-forming, sometimes with solitary culms, rhizomatous. | ||||||||
Culms | 30-150 (200) cm, glabrous or pubescent, sometimes scabrous. |
(10)20-41 cm, smooth, glabrous throughout or pubescent near the inflorescence, bases often geniculate. |
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Sheaths | closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins; collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous; ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent; blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17); abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands; adaxial sclerenchyma sometimes present; girders or pillars present at most veins. |
closed for about 3/4 their length, often splitting with age, coarsely ribbed, shredding into fibers, bases reddish brown, scarious; collars glabrous; ligules 0.1-0.4(0.6) mm; blades 0.3-0.8(1) mm in diameter, conduplicate, green or glaucous, abaxial surfaces glabrous, smooth or scabrous, adaxial surfaces scabrous or puberulent; abaxial sclerenchyma in 5-7(9) small strands; adaxial sclerenchyma absent. |
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Inflorescences | 10-25(30) cm, open, with (1)2(4) branches per node; branches spreading and lax. |
(3)5-12 cm, usually paniculate, sometimes racemose or sub-racemose, compact or open, with 1-2 branches per node; branches stiff or somewhat lax, lower branches with 1-3 spikelets. |
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Spikelets | 8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets. |
pseudoviviparous, varying in length with the stage of vegetative proliferation, the glumes and often 1 or 2 adjacent florets more or less normally developed or only slightly elongated, glabrous or pubescent, the distal florets vegetative. |
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Glumes | lanceolate, glabrous or sparsely scabrous at the apices; lower glumes (4)4.5-6.7(8) mm; upper glumes (5)6-10 mm; lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm; paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally; anthers (3)4-7.5(8.5) mm; ovary apices densely pubescent. |
more or less normally developed, ovate to lanceolate; lower glumes (2.5)3-3.5(5.5); upper glumes 3.5-4.5(6.5); lowest lemma in each spikelet usually normally developed, acute, unawned, usually without reproductive structures or the structures abortive; subsequent lemmas modified into leafy bracts; paleas usually absent, shorter than the lemmas if present, intercostal region puberulent distally; anthers usually aborted, when present 1.5-2.3(3) mm; ovaries rarely present, apices glabrous. |
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2n | = 56. |
= 49, 50, 63. |
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Festuca californica |
Festuca prolifera |
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Distribution |
CA; OR
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Discussion | Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Festuca prolifera is often abundant, and may be a dominant component in some habitats. The leafy bulbils or plantlets sometimes root when the top-heavy inflorescence is bent to the ground. Festuca prolifera has two varieties: Festuca prolifera (Piper) Fernald var. prolifera, with glabrous lemmas; and Festuca prolifera var. lasiolepis Fernald, with pubescent lemmas. Festuca prolifera var. prolifera grows in arctic, alpine, or boreal rocky areas, in calcareous, basic or neutral soils, and is found in the James Bay area, Ungava Bay, western Newfoundland, Cape Breton, the Gaspe Peninsula, the White Mountains (New Hampshire), and Katahdin (Maine). Festuca prolifera var. lasiolepis is found in moist, sandy riverbanks, lake shores, rocky areas, and cliffs, often on limestone, from the southeastern Northwest Territories to northern Quebec, Anticosti Island, and western Newfoundland. Proliferous plants from southern Greenland with extravaginal shoots, named F. villosa-vivipara (Rosenv.) E.B. Alexeev, are similar to F. prolifera, but appear to be hybrids between F. rubra and F. frederikseniae (see under F. frederikseniae, p. 436). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 410. | FNA vol. 24, p. 419. | ||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae | Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca | ||||||||
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Subordinate taxa | ||||||||||
Synonyms | F. rubra var. prolifera, F. rubra subsp. prolifera, F. prolifera var. lasiolepis | |||||||||
Name authority | Vasey | (Piper) Fernald | ||||||||
Web links |