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California fescue

boreal fescue, northern fescue

Habit Plants densely cespitose, without rhizomes. Plants densely cespitose, without rhizomes.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

5-15(20) cm, up to twice as tall as the vegetative shoot leaves, usually erect, sometimes semi-prostrate, glabrous, smooth.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for about 1/2 their length, glabrous, persistent;

collars glabrous;

ligules 0.1-0.5 mm;

blades 0.5-1 mm in diameter, conduplicate, often curved or somewhat falcate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces scabrous, veins 5-7, ribs 3-5;

abaxial sclerenchyma in 3-7 strands, usually less than twice as wide as high;

adaxial sclerenchyma absent.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

1-2(2.5) cm, contracted, usually panicles, sometimes racemes, with 1-2 branches per node;

branches erect, lower branches usually with 1-2 spikelets, sometimes more.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

(3)4-5.5(7) mm, with 3-4(6) florets.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

exceeded by the upper florets, ovate to ovate-lanceolate, mostly glabrous and smooth, sometimes scabrous distally;

lower glumes 1-3.5 mm;

upper glumes 2.2-3.2 mm;

lemmas 2.9-3.5(4.4) mm, ovate, apices scabrous and minutely bidentate, awns (0.5)1.4-2(3) mm, usually slightly subterminal, curved or slightly twisted;

paleas about as long as or slightly longer than the lemmas, intercostal region smooth or scabrous distally;

anthers 0.4-0.8(1.1) mm;

ovary apices glabrous.

Flag

leaf sheaths usually somewhat inflated;

flag leaf blades 0.5-5(8) mm.

2n

= 56.

= 28.

Festuca californica

Festuca hyperborea

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
NF; NT; NU; QC; YT; Greenland
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca hyperborea is a high arctic species that grows from Banks Island in the Canadian Arctic east to Greenland and south to Quebec. It differs from F. brachyphylla (p. 428) in its semi-prostrate habit, the loose sheaths and short blades of its flag leaves, the more pronounced ribs in its lower leaf blades, and its subterminal awn. It differs from F. edlundiae (see next) in having flag leaf blades shorter than 5 mm and smaller spikelets. It has frequently been included in F. ovina (p. 422).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
Source FNA vol. 24, p. 410. FNA vol. 24, p. 432.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
Name authority Vasey Holmen ex Fred.
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