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California fescue

intermountain fescue, oil shale fescue, open fescue

Habit Plants densely cespitose, without rhizomes. Plants loosely or densely cespitose, without rhizomes.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

20-40(50) cm, erect or somewhat geniculate at the base, densely scabrous or pubescent below the inflorescence;

nodes usually not exposed, culms often breaking at the upper nodes at maturity.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for less than 1/2 their length, glabrous, persistent or slowly shredding into fibers;

collars glabrous;

ligules 0.2-0.5 mm;

blades (1)1.2-2.5(3) mm wide, persistent, loosely conduplicate, convolute, or flat, abaxial surfaces glabrous, adaxial surfaces with stiff hairs, veins 7-13, ribs (6)7-13;

abaxial sclerenchyma in strands opposite most of the veins, about as wide as the veins;

adaxial sclerenchyma often present;

pillars or girders sometimes present at the major veins.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

6-12 cm, open, with 2-4 branches per node;

branches stiffly divaricate, densely scabrous-ciliate on the angles, lower branches with 2+ spikelets;

pedicels stiffly hairy.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

5.5-8 mm, with 2(3) florets.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

exceeded by the upper florets, lanceolate-acuminate, sparsely scabrous to puberulent;

lower glumes 3.5-5 mm, distinctly shorter than the adjacent lemmas;

upper glumes 5-7 mm;

lemmas 5-7 mm, chartaceous, scabrous or puberulent, minutely bidentate, awned, awns 1.5-3 mm, subterminal;

paleas about as long as or slightly longer than the lemmas, intercostal region scabrous or puberulent distally;

anthers 1.5-2.5 mm;

ovary apices pubescent.

2n

= 56.

= 28.

Festuca californica

Festuca dasyclada

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; UT
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca dasyclada grows on rocky slopes in open forests and shrublands of western Colorado and central and southern Utah. For many years it was known only from the type collection. When the seeds are mature, the panicles break off the culms and are blown over the ground like a tumbleweed, shedding seeds as they travel. This and other unusual features, such as the divaricate branching pattern and hairy pedicels, prompted WA. Weber to place it in the monotypic genus Argillochloa WA. Weber.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
Source FNA vol. 24, p. 410. FNA vol. 24, p. 443.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
Synonyms Argillochloa dasyclada
Name authority Vasey Hack, ex Beal
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