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California fescue

alpine fescue, fétuque à feuilles courtes

Habit Plants densely cespitose, without rhizomes. Plants densely or loosely cespitose, without rhizomes.
Culms

30-150 (200) cm, glabrous or pubescent, sometimes scabrous.

(5)8-35(55) cm, erect, usually smooth and glabrous, sometimes sparsely scabrous or puberulent near the inflorescence.

Sheaths

closed for less than 1/3 their length, persistent, glabrous or pilose, smooth or scabrous, sometimes scabrous or pilose only distally or on the distal margins;

collars usually densely pubescent or with a few hairs at the margins, sometimes glabrous;

ligules 0.2-5 mm, usually ciliate, abaxial surfaces puberulent;

blades 1-6.5 mm wide, conduplicate, convolute, or flat, 0.5-2(2.5) mm in diameter when convolute, deciduous, abaxial surfaces scabrous or smooth, glabrous or the bases pubescent, adaxial surfaces puberulent to pubescent, veins 9-15(17), ribs (3)5-15(17);

abaxial sclerenchyma forming more or less continuous bands, sometimes reduced to small strands;

adaxial sclerenchyma sometimes present;

girders or pillars present at most veins.

closed for about 1/2 their length, smooth or scabrous, persistent or slowly shredding into fibers;

collars glabrous;

ligules 0.1-0.4 mm;

blades (0.3)0.5-1(1.2) mm in diameter, conduplicate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces scabrous, veins (3)5-7, ribs 3-5;

abaxial sclerenchyma in 3-7(9) narrow strands, usually less than twice as wide as high;

adaxial sclerenchyma absent;

flag leaf sheaths not inflated, more or less tightly enclosing the culms;

flag leaf blades (0.3)1-2.5(3) cm.

Inflorescences

10-25(30) cm, open, with (1)2(4) branches per node;

branches spreading and lax.

1.5-4(5.5) cm, contracted, usually panicles, very rarely racemes, with 1-2 branches per node;

branches usually erect, sometimes spreading at anthesis, lower branches with 2+ spikelets.

Spikelets

8-18(20) mm, borne towards the ends of the branches, with 3-6(8) florets.

3.5-7(8.5) mm, with 2-4(6) florets.

Glumes

lanceolate, glabrous or sparsely scabrous at the apices;

lower glumes (4)4.5-6.7(8) mm;

upper glumes (5)6-10 mm;

lemmas (7)7.5-11 mm, lanceolate, scabrous, puberulent, sometimes minutely bidentate, acute, usually awned, rarely unawned, awns (1)2-3(4) mm;

paleas shorter than to longer than the lemmas, pubescent or glabrous on the margins, intercostal region usually puberulent distally;

anthers (3)4-7.5(8.5) mm;

ovary apices densely pubescent.

exceeded by the upper florets, ovate-lanceolate, usually glabrous and smooth, sometimes scabrous distally;

lower glumes (1.2)1.8-3(3.5) mm;

upper glumes (2.4)2.6-4(4.6) mm;

lemmas 2.5-4.5(6) mm, ovate-lanceolate to lanceolate, scabrous towards the apices, awns (0.8)1-3(3.5) mm, terminal;

paleas about as long as the lemmas, intercostal region scabrous or puberulent distally;

anthers (0.5)0.7-1.1(1.3) mm;

ovary apices glabrous.

2n

= 56.

= 28, 42, 44.

Festuca californica

Festuca brachyphylla

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; ME; MN; MT; NM; NV; OR; UT; VT; WA; WY; AB; BC; MB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
[WildflowerSearch map]
[BONAP county map]
Discussion

Festuca californica grows on dry, open slopes and moist streambanks in thickets and open woods, from sea level to 2000 m. Its range extends from Clackamas County, Oregon, to the Sierra Nevada and southern California; it is not known to extend into Mexico. It is the largest species of Festuca in the Flora region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca brachyphylla is a variable, circumpolar, arctic, alpine, and boreal species of open, rocky places. It is palatable to livestock, and is important in some areas as forage for wildlife. The spikelets are usually tinged red to purple by anthocyanin pigments; plants which lack anthocyanins in the spikelets have been named F. brachyphylla f. flavida Polunin. Festuca brachyphylla has frequently been included in F. ovina (p. 422), and it is closely related to F. saximontana (p. 430), F. hyberborea (p. 432), F. edlundiae (p. 432), F. groenlandica (p. 434), and F. minutiflora (p. 434). It may hybridize with F. baffinensis and/or other species to form F. viviparoidea (p. 436).

Three subspecies have been recognized in North America. Festuca brachyphylla subsp. brachyphylla is circumpolar and primarily arctic, subarctic, and boreal, extending southward in the northern Rocky Mountains. The other two subspecies are restricted to alpine regions in the western mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms 30-80(100) cm tall, usually pubescent for more than 5 mm below the nodes; lower sheaths densely retrorsely pubescent; vegetative shoot blades with (3)5-9 ribs, the ribs to about 1/2 as deep as the blade thickness; abaxial sclerenchyma in small strands or forming continuous bands; adaxial sclerenchyma strands present or absent; sclerenchyma pillars rarely formed; girders not developed; spikelets with 3-4(5) florets
subsp. parishii
1. Culms 60-150(200) cm tall, glabrous or pubescent for less than 5 mm below the nodes; lower sheaths glabrous or pubescent, if pubescent then usually not densely retrorsely hairy; vegetative shoot blades with 7-15(17) ribs, the ribs usually more than 1/2 as deep as the blade thickness; abaxial sclerenchyma forming a continuous band; adaxial sclerenchyma in strands; sclerenchyma pillars or girders usually associated with most of the veins; spikelets with (3)4-6(8) florets.
→ 2
2. Ligules 0.2-1(1.2) mm long, ciliate; spikelets (8)13-18(20) mm long
subsp. californica
2. Ligules (1)1.5-5 mm long, ciliate or not; spikelets 8-12(17) mm long
subsp. hitchcockiana
1. Culms usually more than twice as long as the vegetative shoot leaves; spikelets 4.4-7(8.5) mm long; lemmas (3)3.5-4.5(6) mm long; plants boreal, arctic, and alpine in the northern Cordillera
subsp. brachyphylla
1. Culms up to twice as long as the vegetative shoot leaves; spikelets 3.5-5.5 mm long; lemmas (2.5)3.5-4 mm long; plants alpine in the southern cordillera.
→ 2
2. Culms usually twice as long as the vegetative shoot leaves; awns 2-3(3.2) mm long; spikelets 4.4-5.6(7) mm long; lemmas 3-4(4.5) mm long
subsp. coloradensis
2. Culms usually less than twice as long as the vegetative shoot leaves; awns 1-2(2.2) mm long; spikelets 3.5-5(5.5) mm long; lemmas 2.5-4 mm long
subsp. breviculmis
Source FNA vol. 24, p. 410. FNA vol. 24, p. 428.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Breviaristatae Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. californica subsp. californica, F. californica subsp. hitchcockiana, F. californica subsp. parishii
F. brachyphylla subsp. brachyphylla, F. brachyphylla subsp. breviculmis, F. brachyphylla subsp. coloradensis
Synonyms F. ovina var. brevifolia, F. ovina var. brachyphylla
Name authority Vasey Schult. & Schult. f.
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