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alpine fescue, fétuque à feuilles courtes

Frederksen's fescue, fétuque de frederiksen

Habit Plants densely or loosely cespitose, without rhizomes. Plants densely cespitose, without rhizomes.
Culms

(5)8-35(55) cm, erect, usually smooth and glabrous, sometimes sparsely scabrous or puberulent near the inflorescence.

(5)10-35 (45) cm, pubescent near the inflorescence.

Sheaths

closed for about 1/2 their length, smooth or scabrous, persistent or slowly shredding into fibers;

collars glabrous;

ligules 0.1-0.4 mm;

blades (0.3)0.5-1(1.2) mm in diameter, conduplicate, abaxial surfaces smooth or sparsely scabrous, adaxial surfaces scabrous, veins (3)5-7, ribs 3-5;

abaxial sclerenchyma in 3-7(9) narrow strands, usually less than twice as wide as high;

adaxial sclerenchyma absent;

flag leaf sheaths not inflated, more or less tightly enclosing the culms;

flag leaf blades (0.3)1-2.5(3) cm.

closed for about 1/2 their length, glabrous or puberulent, persistent;

collars glabrous;

ligules 0.2-0.5 mm;

blades 0.5-0.8 mm in diameter, conduplicate, abaxial surfaces glabrous, smooth or scabrous, adaxial surfaces scabrous or hirsute, veins (3)5-7, ribs 3-5, 1 distinct and 2-4 indistinct;

abaxial sclerenchyma in 3-7 broad, sometimes confluent strands, covering 1/2 or more of the surface.

Inflorescences

1.5-4(5.5) cm, contracted, usually panicles, very rarely racemes, with 1-2 branches per node;

branches usually erect, sometimes spreading at anthesis, lower branches with 2+ spikelets.

(1.5)2-10 cm, contracted, with 1(2) branches per node;

branches erect, stiff, lower branches with 2+ spikelets.

Spikelets

3.5-7(8.5) mm, with 2-4(6) florets.

pseudoviviparous, varying in length with the stage of vegetative proliferation, the glumes and often 1 or 2 adjacent florets more or less normally developed or only slightly elongated, the distal florets replaced by leafy bracts.

Glumes

exceeded by the upper florets, ovate-lanceolate, usually glabrous and smooth, sometimes scabrous distally;

lower glumes (1.2)1.8-3(3.5) mm;

upper glumes (2.4)2.6-4(4.6) mm;

lemmas 2.5-4.5(6) mm, ovate-lanceolate to lanceolate, scabrous towards the apices, awns (0.8)1-3(3.5) mm, terminal;

paleas about as long as the lemmas, intercostal region scabrous or puberulent distally;

anthers (0.5)0.7-1.1(1.3) mm;

ovary apices glabrous.

ovate-lanceolate, densely puberulent to pubescent throughout;

lower glumes 2-4.5 mm;

upper glumes (2.7)3.8-5.2 mm;

normal lemmas 3.5-5 mm, densely hairy to pubescent, sometimes awned, awns to 0.2 mm;

vegetative bracts unawned, leaflike, sometimes with ligules;

paleas usually reduced or absent, well-formed paleas about as long as the lemmas;

anthers usually poorly developed and the pollen sterile, well-formed anthers to about 2.5 mm;

ovary apices glabrous.

2n

= 28, 42, 44.

= 28.

Festuca brachyphylla

Festuca frederikseniae

Distribution
from FNA
AK; AZ; CA; CO; ID; ME; MN; MT; NM; NV; OR; UT; VT; WA; WY; AB; BC; MB; NL; NS; NT; NU; ON; QC; SK; YT; Greenland
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Discussion

Festuca brachyphylla is a variable, circumpolar, arctic, alpine, and boreal species of open, rocky places. It is palatable to livestock, and is important in some areas as forage for wildlife. The spikelets are usually tinged red to purple by anthocyanin pigments; plants which lack anthocyanins in the spikelets have been named F. brachyphylla f. flavida Polunin. Festuca brachyphylla has frequently been included in F. ovina (p. 422), and it is closely related to F. saximontana (p. 430), F. hyberborea (p. 432), F. edlundiae (p. 432), F. groenlandica (p. 434), and F. minutiflora (p. 434). It may hybridize with F. baffinensis and/or other species to form F. viviparoidea (p. 436).

Three subspecies have been recognized in North America. Festuca brachyphylla subsp. brachyphylla is circumpolar and primarily arctic, subarctic, and boreal, extending southward in the northern Rocky Mountains. The other two subspecies are restricted to alpine regions in the western mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Festuca frederikseniae grows on cliffs, rocky or sandy barrens, and alpine regions in southern Quebec (Mingan and Anticosti islands), Newfoundland, southern Labrador, and southern Greenland. It differs from F. vivipara (L.) Sm. of northern Europe and Asia in having densely pubescent spikelet bracts and fascicles, and an interrupted rather than continuous band of blade sclerenchyma. Frederiksen (1981) reported that F. vivipara occurs in southeastern Greenland, overlapping the range of F. frederikseniae and extending as far north as the southerly occurrences of F. viviparoidea subsp. viviparoidea; her paper should be consulted when trying to distinguish the complex pseudoviviparous fescues of Greenland.

In Iceland and southern Greenland, putative hybrids between Festuca frederikseniae or F. vivipara and F. rubra (p. 412) have been reported, and named F. villosa-vivipara (Rosenv.) E.B. Alexeev. These plants are highly variable but, unlike F. frederikseniae, produce extravaginal shoots, have closed sheaths, and have blades about 1 mm wide, with 7-9 small strands of abaxial sclerenchyma. Such hybrids can be expected within the range of F. frederikseniae in North America.

Festuca frederikseniae has frequently been included in F. ovina (p. 422).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Culms usually more than twice as long as the vegetative shoot leaves; spikelets 4.4-7(8.5) mm long; lemmas (3)3.5-4.5(6) mm long; plants boreal, arctic, and alpine in the northern Cordillera
subsp. brachyphylla
1. Culms up to twice as long as the vegetative shoot leaves; spikelets 3.5-5.5 mm long; lemmas (2.5)3.5-4 mm long; plants alpine in the southern cordillera.
→ 2
2. Culms usually twice as long as the vegetative shoot leaves; awns 2-3(3.2) mm long; spikelets 4.4-5.6(7) mm long; lemmas 3-4(4.5) mm long
subsp. coloradensis
2. Culms usually less than twice as long as the vegetative shoot leaves; awns 1-2(2.2) mm long; spikelets 3.5-5(5.5) mm long; lemmas 2.5-4 mm long
subsp. breviculmis
Source FNA vol. 24, p. 428. FNA vol. 24, p. 436.
Parent taxa Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca Poaceae > subfam. Pooideae > tribe Poeae > Festuca > subg. Festuca > sect. Festuca
Sibling taxa
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. frederikseniae, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
F. altaica, F. amethystina, F. arizonica, F. auriculata, F. baffinensis, F. brachyphylla, F. brevissima, F. californica, F. calligera, F. campestris, F. dasyclada, F. earlei, F. edlundiae, F. elmeri, F. filiformis, F. glauca, F. groenlandica, F. hallii, F. heterophylla, F. hyperborea, F. idahoensis, F. lenensis, F. ligulata, F. minutiflora, F. occidentalis, F. ovina, F. paradoxa, F. prolifera, F. pseudovivipara, F. roemeri, F. rubra, F. saximontana, F. sororia, F. subulata, F. subuliflora, F. subverticillata, F. thurberi, F. trachyphylla, F. valesiaca, F. versuta, F. viridula, F. viviparoidea, F. washingtonica
Subordinate taxa
F. brachyphylla subsp. brachyphylla, F. brachyphylla subsp. breviculmis, F. brachyphylla subsp. coloradensis
Synonyms F. ovina var. brevifolia, F. ovina var. brachyphylla F. vivipara subsp. hirsuta
Name authority Schult. & Schult. f. E.B. Alexeev
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