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fagonis, sticky fagonbush, sticky fagonia

caltrop family, creosote bush family

Habit Herbs, subshrubs, shrubs, or trees, annual or perennial, branching usually divaricate, growth sympodial, nodes angled or swollen, evergreen [deciduous], synoecious [dioecious].
Stems

± prostrate, radiating from plant base, sparsely branched, dark green, not noticeably slender, ultimate branches densely short-stipitate- to subsessile-glandular, glands globular (drying to cup-shaped), golden (making branches appear yellowish from a distance), 1 mm diam.;

older stems glabrate basally, not scabrous.

Leaves

(1–)3-foliolate;

stipules straight, spreading to slightly reflexed, stout, linear-subulate, 3–16 mm, about as long as petioles, glandular to glabrate;

petiole 2–16 mm, glandular to glabrate;

leaflets ovate to elliptic, slightly obovate, or linear and terete, glandular, becoming glabrate, ± as long as or longer than petiole, apex spinulose, often fleeting, terminal to 26 × 10 mm, laterals to 20 × 7 mm, shorter and narrower than terminal, one or both commonly caducous.

opposite or fascicled [alternate or on short lateral branches], palmately or even- [odd-]pinnately compound [simple];

stipules present;

petiole present [absent];

blade often fleshy or coriaceous, margins entire;

venation pinnate.

Inflorescences

pseudoaxillary [terminal], flowers solitary or in 2-flowered clusters [cymes].

Pedicels

1–7 mm, glandular.

Flowers

to 1.5 cm diam.;

sepals green to purple, ovate-lanceolate, 2–4 × 1–1.5 mm, apiculate, glandular to glabrate;

petals light to dark red-purple, 5–8 × 2.5–5.5 mm;

stamen filaments 3.5–5 mm;

ovary 2 mm, glandular, hairy;

style 2–3 mm.

bisexual [unisexual], usually regular, sometimes slightly irregular;

perianth and androecium hypogynous;

hypanthium absent;

sepals 4–5, usually distinct, rarely connate basally;

petals 4–5, distinct [rarely connate basally], often clawed, sometimes twisted;

nectary usually present, extrastaminal and/or intrastaminal, rarely absent;

stamens [5–](8–)10 in 2 whorls, outer usually opposite petals, often alternately unequal in length or sterile, distinct, free or adnate to petal bases, inserted on or proximal to nectary, frequently glandular or appendaged at base;

anthers dehiscing by longitudinal slits;

pistil 1, (2–)5-carpellate, ovary superior, (2–)5–10-locular;

placentation axile [basal];

ovules (1–)2–10 per locule, anatropous;

style 1;

stigma 1.

Fruits

capsules, dehiscence septicidal or loculicidal, or schizocarps splitting into 5 or 10 mericarps.

Capsules

3.5–5.5 × 4–5 mm, puberulent, usually glandular;

style 1.5–4 mm, not or barely wider at base.

Seeds

1–5(–10) per locule.

Perennial

herbs or subshrubs, to 0.6 m, to 1 m diam.

Fagonia pachyacantha

Zygophyllaceae

Phenology Flowering Nov–May.
Habitat Flat, sandy or rocky desert habitats.
Elevation 0–500 m. (0–1600 ft.)
Distribution
from FNA
AZ; CA; Mexico (Baja California, Baja California Sur, Sonora)
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; Central America; South America; West Indies; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia; mostly tropical or subtropical regions; mainly in arid and semiarid areas
[BONAP county map]
Discussion

Fagonia pachyacantha is found only in the Sonoran Desert.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 27, species ca. 240 (6 genera, 15 species in the flora).

Zygophyllaceae are most closely related to Krameriaceae and the two families make up the isolated order Zygophyllales (Angiosperm Phylogeny Group 2003). A number grow in saline soils. Three species are cultivated in milder winter areas of the southeastern United States for their colorful flowers: the South American Bulnesia arborea (Jacquin) Engler and B. sarmientoi Lorentz ex Grisebach, both verawood, and the Caribbean Guaiacum officinale Linnaeus, lignum vitae. Guaiacum coulteri A. Gray, guayacán, from western Mexico and Guatemala, is grown in southern Arizona. Peganum, often placed in the Zygophyllaceae, is now recognized to be a member of the unrelated Nitrariaceae (Angiosperm Phylogeny Group).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflets 2.
→ 2
2. Leaflets distinct.
Zygophyllum
2. Leaflets connate basally, leaves appearing simple and 2-lobed.
Larrea
1. Leaflets (1–)3 or (4–)6–16(–20).
→ 3
3. Leaves palmately compound; leaflets (1–)3, apex spinose or spinulose.
Fagonia
3. Leaves pinnately compound; leaflets (4–)6–16(–20), apex not spinose.
→ 4
4. Trees or shrubs; petals usually blue to purple, rarely white; fruits capsules, 2–5-lobed.
Guaiacum
4. Herbs; petals white or yellow to bright orange, bases sometimes green or red; fruits schizocarps, 5-angled or 10-lobed.
→ 5
5. Ovaries 5-lobed, 5-locular; fruits 5-angled, spiny, breaking into 5 mericarps (rarely fewer); petals yellow, base darker; nectary 10 glands in 2 whorls.
Tribulus
5. Ovaries 10-lobed, 10-locular; fruits 10-lobed, not spiny, breaking into 10 mericarps (sometimes fewer); petals white to bright orange, base white to bright orange or green to red; nectary 5 glands at bases of filaments opposite petals.
Kallstroemia
Source FNA vol. 12, p. 31. FNA vol. 12, p. 28. Author: Duncan M. Porter.
Parent taxa Zygophyllaceae > Fagonia
Sibling taxa
F. laevis, F. longipes
Subordinate taxa
Fagonia, Guaiacum, Kallstroemia, Larrea, Tribulus, Zygophyllum
Synonyms F. californica var. glutinosa
Name authority Rydberg: in N. L. Britton et al., N. Amer. Fl. 25: 105. (1910) R. Brown
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