Fagonia pachyacantha |
Zygophyllaceae |
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fagonis, sticky fagonbush, sticky fagonia |
caltrop family, creosote bush family |
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Habit | Herbs, subshrubs, shrubs, or trees, annual or perennial, branching usually divaricate, growth sympodial, nodes angled or swollen, evergreen [deciduous], synoecious [dioecious]. | |||||||||||||||||||||
Stems | ± prostrate, radiating from plant base, sparsely branched, dark green, not noticeably slender, ultimate branches densely short-stipitate- to subsessile-glandular, glands globular (drying to cup-shaped), golden (making branches appear yellowish from a distance), 1 mm diam.; older stems glabrate basally, not scabrous. |
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Leaves | (1–)3-foliolate; stipules straight, spreading to slightly reflexed, stout, linear-subulate, 3–16 mm, about as long as petioles, glandular to glabrate; petiole 2–16 mm, glandular to glabrate; leaflets ovate to elliptic, slightly obovate, or linear and terete, glandular, becoming glabrate, ± as long as or longer than petiole, apex spinulose, often fleeting, terminal to 26 × 10 mm, laterals to 20 × 7 mm, shorter and narrower than terminal, one or both commonly caducous. |
opposite or fascicled [alternate or on short lateral branches], palmately or even- [odd-]pinnately compound [simple]; stipules present; petiole present [absent]; blade often fleshy or coriaceous, margins entire; venation pinnate. |
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Inflorescences | pseudoaxillary [terminal], flowers solitary or in 2-flowered clusters [cymes]. |
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Pedicels | 1–7 mm, glandular. |
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Flowers | to 1.5 cm diam.; sepals green to purple, ovate-lanceolate, 2–4 × 1–1.5 mm, apiculate, glandular to glabrate; petals light to dark red-purple, 5–8 × 2.5–5.5 mm; stamen filaments 3.5–5 mm; ovary 2 mm, glandular, hairy; style 2–3 mm. |
bisexual [unisexual], usually regular, sometimes slightly irregular; perianth and androecium hypogynous; hypanthium absent; sepals 4–5, usually distinct, rarely connate basally; petals 4–5, distinct [rarely connate basally], often clawed, sometimes twisted; nectary usually present, extrastaminal and/or intrastaminal, rarely absent; stamens [5–](8–)10 in 2 whorls, outer usually opposite petals, often alternately unequal in length or sterile, distinct, free or adnate to petal bases, inserted on or proximal to nectary, frequently glandular or appendaged at base; anthers dehiscing by longitudinal slits; pistil 1, (2–)5-carpellate, ovary superior, (2–)5–10-locular; placentation axile [basal]; ovules (1–)2–10 per locule, anatropous; style 1; stigma 1. |
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Fruits | capsules, dehiscence septicidal or loculicidal, or schizocarps splitting into 5 or 10 mericarps. |
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Capsules | 3.5–5.5 × 4–5 mm, puberulent, usually glandular; style 1.5–4 mm, not or barely wider at base. |
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Seeds | 1–5(–10) per locule. |
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Perennial | herbs or subshrubs, to 0.6 m, to 1 m diam. |
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Fagonia pachyacantha |
Zygophyllaceae |
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Phenology | Flowering Nov–May. | |||||||||||||||||||||
Habitat | Flat, sandy or rocky desert habitats. | |||||||||||||||||||||
Elevation | 0–500 m. (0–1600 ft.) | |||||||||||||||||||||
Distribution |
AZ; CA; Mexico (Baja California, Baja California Sur, Sonora)
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North America; Mexico; Central America; South America; West Indies; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia; mostly tropical or subtropical regions; mainly in arid and semiarid areas |
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Discussion | Fagonia pachyacantha is found only in the Sonoran Desert. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 27, species ca. 240 (6 genera, 15 species in the flora). Zygophyllaceae are most closely related to Krameriaceae and the two families make up the isolated order Zygophyllales (Angiosperm Phylogeny Group 2003). A number grow in saline soils. Three species are cultivated in milder winter areas of the southeastern United States for their colorful flowers: the South American Bulnesia arborea (Jacquin) Engler and B. sarmientoi Lorentz ex Grisebach, both verawood, and the Caribbean Guaiacum officinale Linnaeus, lignum vitae. Guaiacum coulteri A. Gray, guayacán, from western Mexico and Guatemala, is grown in southern Arizona. Peganum, often placed in the Zygophyllaceae, is now recognized to be a member of the unrelated Nitrariaceae (Angiosperm Phylogeny Group). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 31. | FNA vol. 12, p. 28. | ||||||||||||||||||||
Parent taxa | Zygophyllaceae > Fagonia | |||||||||||||||||||||
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Synonyms | F. californica var. glutinosa | |||||||||||||||||||||
Name authority | Rydberg: in N. L. Britton et al., N. Amer. Fl. 25: 105. (1910) | R. Brown | ||||||||||||||||||||
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