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drug eyebright, euphraise dressée, stiff eyebright, strict eyebright, Tartary eyebright

arctic eyebright

Stems

branched, rarely simple, to 30 cm;

branches 1–5 pairs, from middle and distal cauline nodes;

cauline internode length 0.5–2 times subtending leaves.

simple or branched, to 25(–35) cm;

branches 1–5(or 6) pairs, from proximal, middle, or distal cauline nodes;

cauline internode lengths 2–4 times subtending leaves.

Leaves

blade narrowly oblong to ovate-oblong, 2–9(–12) mm, margins serrate, teeth 1–3 pairs, apices subacute to aristate.

blade usually broadly ovate, 3–16 mm, margins serrate, teeth 1–6 pairs, apices acute.

Inflorescences

beginning at node (5–)7–9(–11);

bracts green, sometimes purple adaxially, broader than leaves, oblong to ovate, length not more than 2 times width, 5–10 mm, surfaces glabrous or sparsely to densely hirsute and hairs eglandular, sometimes glandular-pubescent and hairs glandular, stalks 1- or 2-celled, 0.1–0.2 mm, teeth 4 or 5 pairs, longer than wide, apices acute, sometimes aristate.

beginning at node 3–5;

bracts green or purplish adaxially, broader than leaves, broadly ovate or deltate, length not more than 2 times width, (4–)5–9(–11) mm, surfaces glabrous or setose and hairs eglandular, sometimes pubescent and hairs glandular, stalks 1- or 2-celled, 0.1–0.2 mm, teeth 3–6 pairs, as long as or much longer than wide, apices acute to acuminate.

Flowers

corolla white, usually suffused with lilac, 8–10 mm, abaxial lip exceeding adaxial.

corolla white, sometimes lilac, adaxial lip lilac, 6–11(–13) mm, abaxial lip exceeding adaxial.

Capsules

narrowly oblong, 4.5–6 mm, apex truncate to retuse.

oblong to elliptic or obovate, 4.5–8 mm, apex truncate to retuse.

Euphrasia stricta

Euphrasia arctica

Distribution
from FNA
IL; MA; ME; MI; MN; NH; NY; PA; RI; VT; WI; NB; NS; ON; PE; QC; Europe; Asia (w Siberia) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
ME; BC; NB; NF; NS; ON; PE; QC; Europe; Atlantic Islands (Iceland)
Discussion

Varieties 7 (2 in the flora).

Occurrence of Euphrasia stricta in Alberta is based on the report by G. C. D. Griffiths (2002); no specimen has been located. Griffiths listed E. arctica subsp. borealis as new for Alberta, but both morphological and habitat descriptions fit E. stricta more closely. If true, then this introduced species has crossed the continent; all earlier records are from eastern Canada and New England (S. R. Downie and J. McNeill 1988).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (2 in the flora).

The other three subspecies of Euphrasia arctica occur in Europe (P. F. Yeo 1978).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Bract surfaces glabrous or sparsely to densely hirsute, hairs eglandular.
var. stricta
1. Bract surfaces hairy, hairs glandular.
var. brevipila
1. Stem bases flexuous or decumbent; branches 1 or 2(–5) pairs, usually flexuous; bract surfaces glabrous or hairy, hairs eglandular bristles, teeth not much longer than wide; corollas 7–11(–13) mm.
subsp. arctica
1. Stem bases erect; branches 1–5(or 6) pairs, usually straight; bract surfaces hairy, hairs glandular and eglandular bristles, teeth much longer than wide; corollas 6–8(–10) mm.
subsp. borealis
Source FNA vol. 17, p. 496. FNA vol. 17, p. 495.
Parent taxa Orobanchaceae > Euphrasia Orobanchaceae > Euphrasia
Sibling taxa
E. arctica, E. disjuncta, E. farlowii, E. frigida, E. hudsoniana, E. micrantha, E. mollis, E. nemorosa, E. oakesii, E. ostenfeldii, E. randii, E. salisburgensis, E. subarctica, E. suborbicularis, E. tetraquetra, E. vinacea, E. wettsteinii
E. disjuncta, E. farlowii, E. frigida, E. hudsoniana, E. micrantha, E. mollis, E. nemorosa, E. oakesii, E. ostenfeldii, E. randii, E. salisburgensis, E. stricta, E. subarctica, E. suborbicularis, E. tetraquetra, E. vinacea, E. wettsteinii
Subordinate taxa
E. stricta var. brevipila, E. stricta var. stricta
E. arctica subsp. arctica, E. arctica subsp. borealis
Name authority J. P. Wolff ex J. F. Lehmann: Prim. Lin. Fl. Herbipol., 43. (1809) Lange ex Rostrup: Bot. Tidsskr. 4: 47. (1870)
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