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hairy spurge

perennial sandmat, Terlingua spurge

Habit Herbs, annual or perennial, with slender taproot or thickened, woody rootstock. Herbs, perennial, with strongly thickened, woody rootstock.
Stems

usually erect to ascending, rarely prostrate to decumbent, 10–30 cm, papillate, usually villous, sometimes glabrous.

erect, 7–45 cm, glabrous.

Leaves

opposite;

stipules distinct, filiform, usually undivided, rarely divided into 2–3 segments), 0.3–0.7 mm, glabrous, papillate;

petiole 0.6–1.8 mm, usually villous, rarely glabrous;

blade ovate, 3–12 × 2–10 mm, base asymmetric, rounded to slightly cordate, margins entire or serrulate, apex acute to obtuse, surfaces usually villous, rarely glabrous;

only midvein conspicuous.

opposite;

stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous;

petiole 0.8–2 mm, glabrous;

blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous.

Involucre

campanulate, 0.7–0.9 × 0.6–1 mm, glabrous or pilose;

glands 4, pink, oval, oblong, or trapezoidal, 0.1–0.2 × 0.2 mm;

appendages white to pink, flabellate, oblong, ovate, or nearly rectangular, 0.2–0.4 × 0.2–0.6 mm, distal margin entire.

broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous;

glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm;

appendages absent.

Staminate flowers

10–25.

35–45.

Pistillate flowers

ovary glabrous;

styles 0.3–0.5 mm, 2-fid 1/2 to nearly entire length.

ovary glabrous;

styles 0.7–0.9 mm, 2-fid nearly entire length.

Capsules

oblate-deltoid, cocci often elongated and terminating in an empty portion, 1.5–2 × 2.1–3.1 mm, glabrous;

columella 0.9–1.5 mm.

subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous;

columella 2.2–2.7 mm.

Seeds

gray-brown to red-brown, ovoid-oblong, weakly 4-angled in cross section, 1–1.4 × 0.6–0.8 mm, smooth, faintly rugose, or with inconspicuous transverse ridges.

white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled.

Cyathia

solitary at distal nodes;

peduncles 0–1.8 mm.

solitary at distal nodes or at nodes of short, axillary branches;

peduncle 1.8–3 mm.

Euphorbia villifera

Euphorbia perennans

Phenology Flowering and fruiting early spring–early winter. Flowering and fruiting spring–fall.
Habitat Riparian forests with walnuts and sycamores, juniper woodlands, pine-oak woodlands, mostly on limestone substrates. Desert scrub, on cretaceous and gypseous clay, limestone hills and flats.
Elevation 100–1400 m. (300–4600 ft.) 900–1200 m. (3000–3900 ft.)
Distribution
from FNA
TX; Mexico; Central America
[BONAP county map]
from FNA
TX; Mexico (Chihuahua)
[BONAP county map]
Discussion

Although Euphorbia villifera has been reported from New Mexico (W. C. Martin and C. R. Hutchins 1980), no vouchers to verify its presence there were located. In Texas, E. villifera is known from the Edwards Plateau westward into the trans-Pecos region.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011).

Euphorbia perennans is known in the flora area only from Brewster County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 293. FNA vol. 12, p. 282.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce stanfieldii, C. villifera, E. stanfieldii, E. villifera var. nuda Chamaesyce perennans
Name authority Scheele: Linnaea 22: 153. (1849) (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960)
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