Euphorbia vermiculata |
Euphorbia mesembrianthemifolia |
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hairy sandmat, wormseed sandmat, wormseed spurge |
coastal beach sandmat |
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Habit | Herbs, annual or short-lived perennial, with slender, fibrous taproot. | Subshrubs or shrubs, perennial, with thickened and often woody rootstock. |
Stems | prostrate to ascending or erect, not mat forming, 10–35 cm, usually sparsely to moderately strigillose, pilose, or villous, rarely glabrate hairs sometimes in longitudinal lines. |
erect to ascending, or nearly decumbent in shifting sand, 25–60 cm, glabrous. |
Leaves | opposite; stipules distinct or connate, triangular to narrowly triangular or laciniate into subulate to filiform divisions, 0.6–1.3 mm, glabrous or sparsely pilose; petiole 0.2–0.9 mm, glabrous, villous, or strigillose; blade ovate, oblong, or elliptic, often falcate, 5–18 × 3–9 mm, base asymmetric, one side rounded and other cordate, margins usually serrulate especially in distal 1/2, rarely nearly entire, apex acute or obtuse, surfaces not papillate, sparsely pilose, villous, or sericeous (especially near base), often glabrate (especially older leaves); 3–5-veined from base. |
opposite; stipules connate, forming conspicuous, ligulate or deltate scale, short cleft or fringed, 1–1.8 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to elliptic often folded along midrib, 5–12 × 3–8 mm, ± fleshy, base slightly asymmetric, truncate to cordate, partially obscuring stem, margins entire, apex usually obtuse, rarely acute, surfaces yellowish to dark green, glabrous glaucous; obscurely 3–5-veined at base, pinnate distally, only midvein conspicuous. |
Involucre | obconic, 0.7–1.1 × 0.5–0.8 mm, glabrous; glands (2–3)–4, red to reddish green, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent or white, turning pink with age, flabellate, oblong, circular, or forming narrow lunate border around margin of gland, 0.1–0.3 × 0.2–0.4 mm, distal margin entire or slightly lobed. |
campanulate, 1–1.6 × 0.7–1.2 mm, glabrous; glands 4, brown, usually elliptic, occasionally almost round, 0.2–0.4 × 0.5–0.7 mm, fleshy; appendages white, oblong, rarely rudimentary, 0.2–0.4 × 0.5–0.9 mm, distal margin entire or undulate. |
Staminate flowers | 5–15. |
12–20. |
Pistillate flowers | ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
Capsules | oblate to subglobose, cocci not elongated nor terminating in empty portion, 1.4–1.8 × 1.7–2.1 mm, glabrous; columella 1.1–1.5 mm. |
subglobose subtended by calyxlike structure, 1.5–2 × 2.2–2.8 mm, glabrous; columella 1–1.5 mm. |
Seeds | brown, gray, or almost black, ovoid to oblong, 3–4-angled in cross section, 1.1–1.4 × 0.7–0.8 mm, rugulose and sometimes also with low transverse ridges that do not interrupt abaxial keel. |
ashen, broadly ovoid, angled in cross section, faces plump, convex, 1.2–1.3 × 0.9–1.2 mm, obscurely pitted. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.2–2.5 mm. |
solitary at distal nodes; peduncle 0.5–1 mm. |
Euphorbia vermiculata |
Euphorbia mesembrianthemifolia |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting year-round. |
Habitat | Juniper-oak woodlands, temperate deciduous forests, grasslands, pine forests, oak forests with sycamores, walnuts and alders, often in disturbed areas. | Sandy and rocky shores, associated beach scrub. |
Elevation | 0–2600 m. (0–8500 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; CT; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NM; NY; OH; PA; RI; VA; VT; WI; WV; NB; NS; ON; QC; Mexico; temperate deciduous forests; often in disturbed areas; walnuts and alders; oak forests with sycamores; pine forests; grasslands; Juniper-oak woodlands
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FL; Mexico; West Indies; Bermuda; South America (Colombia, Venezuela) |
Discussion | Euphorbia vermiculata has an interesting disjunct distribution; it ranges from central Mexico to Arizona and New Mexico, and is also present in the northeastern United States and southeastern Canada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia mesembrianthemifolia is found in the flora area along the sandy and rocky shores of southern Florida from Pinellas and Volusia counties southward. It is one of the few members of subg. Chamaesyce in the flora area that is a shrub or subshrub. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 292. | FNA vol. 12, p. 277. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce rothrockii, C. vermiculata, E. rothrockii | Chamaesyce buxifolia, C. mesembrianthemifolia, E. buxifolia |
Name authority | Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2: 206. (1818) | Jacquin: Enum. Syst. Pl., 22. (1760) |
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