Euphorbia vermiculata |
Euphorbia inundata |
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hairy sandmat, wormseed sandmat, wormseed spurge |
Florida pineland spurge |
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Habit | Herbs, annual or short-lived perennial, with slender, fibrous taproot. | Herbs, perennial, with thickened rootstock. | ||||
Stems | prostrate to ascending or erect, not mat forming, 10–35 cm, usually sparsely to moderately strigillose, pilose, or villous, rarely glabrate hairs sometimes in longitudinal lines. |
erect or ascending, 15–40 cm. |
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Leaves | opposite; stipules distinct or connate, triangular to narrowly triangular or laciniate into subulate to filiform divisions, 0.6–1.3 mm, glabrous or sparsely pilose; petiole 0.2–0.9 mm, glabrous, villous, or strigillose; blade ovate, oblong, or elliptic, often falcate, 5–18 × 3–9 mm, base asymmetric, one side rounded and other cordate, margins usually serrulate especially in distal 1/2, rarely nearly entire, apex acute or obtuse, surfaces not papillate, sparsely pilose, villous, or sericeous (especially near base), often glabrate (especially older leaves); 3–5-veined from base. |
petiole absent or indistinct, blade linear to linear-elliptic or lanceolate, (25–)30–60(–115) × 1.5–14(–15) mm, chartaceous, base attenuate, apex acuminate, acute, or rounded and mucronate; only midvein evident. |
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Involucre | obconic, 0.7–1.1 × 0.5–0.8 mm, glabrous; glands (2–3)–4, red to reddish green, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent or white, turning pink with age, flabellate, oblong, circular, or forming narrow lunate border around margin of gland, 0.1–0.3 × 0.2–0.4 mm, distal margin entire or slightly lobed. |
obconic or campanulate, 1.3–3.2 × 1.6–3.6 mm, lobes ovate, oblong, or nearly rectangular, 0.6–1 mm, laciniate-ciliate; glands red to greenish, oblong or nearly circular, 0.8–1 × 1.1–2 mm, distal margins crenulate-erose, undulate, or entire. |
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Staminate flowers | 5–15. |
20–25. |
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Pistillate flowers | ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
gynophore exserted 2.6–6.1 mm, calyxlike lobes triangular to subulate, 0.6–1.4 mm; styles connate 1/8 length, 1.3–2.2 mm. |
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Capsules | oblate to subglobose, cocci not elongated nor terminating in empty portion, 1.4–1.8 × 1.7–2.1 mm, glabrous; columella 1.1–1.5 mm. |
oblate-ovoid, 5.1–6.1 × 6.9–8.6 mm, 3-lobed; columella 3.9–4.7 mm. |
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Seeds | brown, gray, or almost black, ovoid to oblong, 3–4-angled in cross section, 1.1–1.4 × 0.7–0.8 mm, rugulose and sometimes also with low transverse ridges that do not interrupt abaxial keel. |
brown to blackish, ovoid-globose, circular or faintly 3- or 4-angled in cross section, 2.9–3.4 × 2.7–3 mm, smooth, base flattened or rounded, apex rounded or with inconspicuous blunt point. |
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Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.2–2.5 mm. |
peduncle 6–24 mm (often exceeding dichasial bracts). |
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Cyathial | arrangement: terminal pleiochasial branches 2–3, 5–18 cm, 3–8 times 2-branched; pleiochasial bracts linear-lanceolate or narrowly ovate, 25–49 × 5–9 mm, margins entire, apex acute or acuminate; dichasial bracts ovate or lanceolate, 6–21 × 2–6 mm, margins entire, apex acute or acuminate; axillary cymose branches 1–5. |
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Euphorbia vermiculata |
Euphorbia inundata |
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Phenology | Flowering and fruiting spring–fall. | |||||
Habitat | Juniper-oak woodlands, temperate deciduous forests, grasslands, pine forests, oak forests with sycamores, walnuts and alders, often in disturbed areas. | |||||
Elevation | 0–2600 m. (0–8500 ft.) | |||||
Distribution |
AZ; CT; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NM; NY; OH; PA; RI; VA; VT; WI; WV; NB; NS; ON; QC; Mexico; temperate deciduous forests; often in disturbed areas; walnuts and alders; oak forests with sycamores; pine forests; grasslands; Juniper-oak woodlands
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AL; FL; GA; MS |
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Discussion | Euphorbia vermiculata has an interesting disjunct distribution; it ranges from central Mexico to Arizona and New Mexico, and is also present in the northeastern United States and southeastern Canada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia inundata has a disjunct distribution in the southeastern United States. Allopatric, narrow-leaved populations from the west-central Florida peninsula are segregated as var. garrettii (E. L. Bridges and S. L. Orzell 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 292. | FNA vol. 12, p. 314. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Nummulariopsis | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce rothrockii, C. vermiculata, E. rothrockii | Galarhoeus inundatus | ||||
Name authority | Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2: 206. (1818) | Torrey ex Chapman: Fl. South. U.S., 402. (1860) | ||||
Web links |