Euphorbia vermiculata |
Euphorbia crenulata |
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hairy sandmat, wormseed sandmat, wormseed spurge |
Chinese caps, western wood spurge |
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Habit | Herbs, annual or short-lived perennial, with slender, fibrous taproot. | Herbs, usually biennial, occasionally annual, with taproot. |
Stems | prostrate to ascending or erect, not mat forming, 10–35 cm, usually sparsely to moderately strigillose, pilose, or villous, rarely glabrate hairs sometimes in longitudinal lines. |
erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
Leaves | opposite; stipules distinct or connate, triangular to narrowly triangular or laciniate into subulate to filiform divisions, 0.6–1.3 mm, glabrous or sparsely pilose; petiole 0.2–0.9 mm, glabrous, villous, or strigillose; blade ovate, oblong, or elliptic, often falcate, 5–18 × 3–9 mm, base asymmetric, one side rounded and other cordate, margins usually serrulate especially in distal 1/2, rarely nearly entire, apex acute or obtuse, surfaces not papillate, sparsely pilose, villous, or sericeous (especially near base), often glabrate (especially older leaves); 3–5-veined from base. |
petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | obconic, 0.7–1.1 × 0.5–0.8 mm, glabrous; glands (2–3)–4, red to reddish green, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent or white, turning pink with age, flabellate, oblong, circular, or forming narrow lunate border around margin of gland, 0.1–0.3 × 0.2–0.4 mm, distal margin entire or slightly lobed. |
campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
Staminate flowers | 5–15. |
11–18. |
Pistillate flowers | ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
Capsules | oblate to subglobose, cocci not elongated nor terminating in empty portion, 1.4–1.8 × 1.7–2.1 mm, glabrous; columella 1.1–1.5 mm. |
subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
Seeds | brown, gray, or almost black, ovoid to oblong, 3–4-angled in cross section, 1.1–1.4 × 0.7–0.8 mm, rugulose and sometimes also with low transverse ridges that do not interrupt abaxial keel. |
cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.2–2.5 mm. |
peduncle 0–0.5 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
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Euphorbia vermiculata |
Euphorbia crenulata |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting spring–summer. |
Habitat | Juniper-oak woodlands, temperate deciduous forests, grasslands, pine forests, oak forests with sycamores, walnuts and alders, often in disturbed areas. | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. |
Elevation | 0–2600 m. (0–8500 ft.) | 30–1800 m. (100–5900 ft.) |
Distribution |
AZ; CT; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NM; NY; OH; PA; RI; VA; VT; WI; WV; NB; NS; ON; QC; Mexico; temperate deciduous forests; often in disturbed areas; walnuts and alders; oak forests with sycamores; pine forests; grasslands; Juniper-oak woodlands
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CA; CO; NM; OR
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Discussion | Euphorbia vermiculata has an interesting disjunct distribution; it ranges from central Mexico to Arizona and New Mexico, and is also present in the northeastern United States and southeastern Canada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 292. | FNA vol. 12, p. 301. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce rothrockii, C. vermiculata, E. rothrockii | Tithymalus crenulatus |
Name authority | Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2: 206. (1818) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) |
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