Euphorbia vermiculata |
Euphorbia blodgettii |
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hairy sandmat, wormseed sandmat, wormseed spurge |
limestone sandmat |
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Habit | Herbs, annual or short-lived perennial, with slender, fibrous taproot. | Herbs, usually annual, occasionally perennial, with slender to slightly thickened taproot, 3.5 mm diam.. |
Stems | prostrate to ascending or erect, not mat forming, 10–35 cm, usually sparsely to moderately strigillose, pilose, or villous, rarely glabrate hairs sometimes in longitudinal lines. |
prostrate to decumbent, loosely mat-forming, often rooting at nodes, 10–45 cm, glabrous. |
Leaves | opposite; stipules distinct or connate, triangular to narrowly triangular or laciniate into subulate to filiform divisions, 0.6–1.3 mm, glabrous or sparsely pilose; petiole 0.2–0.9 mm, glabrous, villous, or strigillose; blade ovate, oblong, or elliptic, often falcate, 5–18 × 3–9 mm, base asymmetric, one side rounded and other cordate, margins usually serrulate especially in distal 1/2, rarely nearly entire, apex acute or obtuse, surfaces not papillate, sparsely pilose, villous, or sericeous (especially near base), often glabrate (especially older leaves); 3–5-veined from base. |
opposite; stipules distinct, subulate filiform segments (lower side), or connate forming conspicuous, broad deltate scale (upper side), toothed, 0.5–1 mm, glabrous; petiole 0.7–1 mm, glabrous; blade ovate to oblong-elliptic, 4–10 × 2–5 mm, base asymmetric, subcordate to rounded, margins usually entire, occasionally toothed, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces without red blotch, glabrous; 3-veined from base, only midvein conspicuous. |
Involucre | obconic, 0.7–1.1 × 0.5–0.8 mm, glabrous; glands (2–3)–4, red to reddish green, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent or white, turning pink with age, flabellate, oblong, circular, or forming narrow lunate border around margin of gland, 0.1–0.3 × 0.2–0.4 mm, distal margin entire or slightly lobed. |
turbinate to campanulate, 0.5–0.6 × 0.4–0.6 mm, glabrous; glands 4, red, slightly concave, elliptic-oblong, 0.1 × 0.1–0.3 mm; appendages white to pink, unequal, pair near sinus lunate to oblong, 0.1–0.2 × 0.1–0.3 mm, distal margin entire, crenulate, or irregularly sinuate, other pair sometimes rudimentary, 0–0.1 × 0.1–0.3 mm, distal margin crenulate or entire. |
Staminate flowers | 5–15. |
8–12. |
Pistillate flowers | ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
Capsules | oblate to subglobose, cocci not elongated nor terminating in empty portion, 1.4–1.8 × 1.7–2.1 mm, glabrous; columella 1.1–1.5 mm. |
broadly ovoid, 1.2–1.7 × 1.4–1.9 mm, glabrous; columella 1.2–1.6 mm. |
Seeds | brown, gray, or almost black, ovoid to oblong, 3–4-angled in cross section, 1.1–1.4 × 0.7–0.8 mm, rugulose and sometimes also with low transverse ridges that do not interrupt abaxial keel. |
gray to reddish brown, oblong-ovoid, 4-angled in cross section, 0.9–1.1 × 0.5–0.6 mm, flat or obscurely wrinkled. |
Cyathia | solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.2–2.5 mm. |
solitary or in small, cymose clusters at distal nodes of stem or on congested, axillary branches; peduncle 0.4–0.6 mm. |
Euphorbia vermiculata |
Euphorbia blodgettii |
|
Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting year-round. |
Habitat | Juniper-oak woodlands, temperate deciduous forests, grasslands, pine forests, oak forests with sycamores, walnuts and alders, often in disturbed areas. | Coastal sand dunes and disturbed upland sandy areas. |
Elevation | 0–2600 m. (0–8500 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
AZ; CT; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NM; NY; OH; PA; RI; VA; VT; WI; WV; NB; NS; ON; QC; Mexico; temperate deciduous forests; often in disturbed areas; walnuts and alders; oak forests with sycamores; pine forests; grasslands; Juniper-oak woodlands
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FL |
Discussion | Euphorbia vermiculata has an interesting disjunct distribution; it ranges from central Mexico to Arizona and New Mexico, and is also present in the northeastern United States and southeastern Canada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia blodgettii is found only in peninsular Florida. It is closely related to E. garberi, E. porteriana, and E. serpens (Y. Yang and P. E. Berry 2011). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 292. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce rothrockii, C. vermiculata, E. rothrockii | Chamaesyce blodgettii, C. nashii |
Name authority | Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2: 206. (1818) | Engelmann ex Hitchcock: Rep. (Annual) Missouri Bot. Gard. 4: 126, plate 13. (1893) |
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