Euphorbia thymifolia |
Euphorbia tetrapora |
|
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gulf sandmat |
weak spurge |
|
Habit | Herbs, annual or perennial, with taproot. | Herbs, annual, with taproot. |
Stems | prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose. |
erect, unbranched, 7–20 cm, glabrous. |
Leaves | opposite; stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose; petiole 0.5–1 mm, sparsely strigose-tomentulose; blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally. |
petiole 1–2 mm, reduced distally; blade spatulate-cuneate, 8–10 × 4–5 mm (greatly reduced in size proximally), base cuneate, margins entire, apex rounded to emarginate or obcordate, surfaces glabrous; venation pinnate. |
Involucre | broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose; glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate. |
campanulate, 0.8–1.1 × 0.7–1 mm, glabrous; glands 4, elliptic to trapezoidal, 0.3–0.6 × 0.6–1.2 mm; horns divergent, 0.5–1 mm. |
Staminate flowers | 3–5. |
10–15. |
Pistillate flowers | ovary densely strigose; styles 0.4–0.5 mm, 2-fid 1/2 length. |
ovary smooth, glabrous; styles 0.6–1 mm, 2-fid. |
Capsules | conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose; columella 0.6–1 mm. |
depressed-globose, 1.8–2.2 × 2.2–2.9 mm, slightly lobed; cocci rounded to slightly flattened, smooth, glabrous; columella 1.5–1.8 mm. |
Seeds | white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate. |
reddish brown to brown, often glaucous, oblong, 1.3–1.4 × 0.8–0.9 mm, abaxial faces with 15–20 shallow pits or almost smooth, adaxial faces with 4–6 large shallow pits or irregular oblong grooves; caruncle conic, hat-shaped, 0.3–0.4 × 0.4–0.6 mm. |
Cyathia | usually in small, cymose clusters on congested, axillary branches; peduncle (0–)0.1–0.3 mm. |
peduncle 0.2–0.6 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 1–3(–4) times 2-branched; pleiochasial bracts obovate, similar in size to distal leaves; dichasial bracts distinct or basally subconnate, not imbricate, triangular-ovate, base truncate or cordate, margins entire, apex mucronate; axillary cymose branches 1–4. |
|
Euphorbia thymifolia |
Euphorbia tetrapora |
|
Phenology | Flowering and fruiting midsummer–early fall. | Flowering and fruiting spring. |
Habitat | Disturbed areas, often near salt water. | Sandy soils, dry open woods. |
Elevation | 0–20 m. (0–100 ft.) | 0–300 m. (0–1000 ft.) |
Distribution |
FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia] |
LA; OK; TX |
Discussion | Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia tetrapora is endemic to a portion of the western Gulf coastal plain. D. S. Correll and M. C. Johnston (1970) included Alabama and Georgia in the distribution of this species as well, probably due to Engelmann’s citation of a Georgia specimen from the herbarium of Samuel Boykin. Whether the Boykin specimen came from Georgia, where Boykin was based, is unclear. Because no records to support its occurrence in the eastern Gulf coastal plain (Alabama or Georgia) have been found, those states are here excluded from the distribution of E. tetrapora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 291. | FNA vol. 12, p. 311. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce thymifolia | Tithymalus tetraporus |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 191. (1859) |
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