FNA: Euphorbia thymifolia vs. Euphorbia simulans

	Euphorbia thymifolia	Euphorbia simulans
	gulf sandmat	mimicking sandmat, similar spurge
Habit	Herbs, annual or perennial, with taproot.	Herbs, annual or short-lived perennial, with usually slender, occasionally slightly thickened, rootstock.
Stems	prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose.	prostrate to reclining, 5–40 cm, glabrous.
Leaves	opposite; stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose; petiole 0.5–1 mm, sparsely strigose-tomentulose; blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally.	opposite; stipules distinct, subulate, 0.5–0.7 mm, glabrous; petiole 0.7–1.3 mm, glabrous; blade orbiculate, oval, to shortly oblong, 1–3.2 × 1.5–5 mm, base subsymmetric, rounded, margins entire, apex usually rounded, occasionally emarginate, surfaces glabrous; venation obscure, only midvein conspicuous.
Involucre	broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose; glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate.	turbinate to campanulate, 0.8–1.2 × 0.7–1 mm, glabrous; glands 4, red to purple, slightly concave, elliptic, 0.2–0.3 × 0.4–0.5 mm; appendages absent.
Staminate flowers	3–5.	15–36.
Pistillate flowers	ovary densely strigose; styles 0.4–0.5 mm, 2-fid 1/2 length.	ovary glabrous; styles 0.2–0.3 mm, 2-fid nearly entire length.
Capsules	conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose; columella 0.6–1 mm.	broadly ovoid, 1.3–1.8 × 1.5–2 mm, glabrous; columella 1.1–1.4 mm.
Seeds	white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate.	whitish, reddish brown beneath coat, oblong, 4-angled in cross section, 1.5–2 × 1.3–1.8 mm, with 5–7 faint transverse ridges or wrinkles.
Cyathia	usually in small, cymose clusters on congested, axillary branches; peduncle (0–)0.1–0.3 mm.	solitary at distal nodes; peduncle 0.3–0.7 mm.
2n		= 28.

	Euphorbia thymifolia	Euphorbia simulans
Phenology	Flowering and fruiting midsummer–early fall.	Flowering and fruiting year-round.
Habitat	Disturbed areas, often near salt water.	Desert scrub, mountains, hills, canyons, arroyos, flats, roadsides, clay, sandy, gravelly, and rocky soils.
Elevation	0–20 m. (0–100 ft.)	600–1300 m. (2000–4300 ft.)
Distribution	FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia] [BONAP county map]	TX; Mexico (Chihuahua, Coahuila) [BONAP county map]
Discussion	Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Euphorbia simulans, which in the flora area is known only from Brewster, Hudspeth, and Presidio counties, is difficult to distinguish in the field from the sympatric E. theriaca var. theriaca, because they are mainly distinguished by seed morphology. The latter has smaller seeds with (two or) three (or four) prominent transverse ridges, whereas E. simulans has larger seeds that are slightly wrinkled. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 12, p. 291.	FNA vol. 12, p. 289.
Parent taxa	Euphorbiaceae > Euphorbia > sect. Anisophyllum	Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms	Chamaesyce thymifolia	E. polycarpa var. simulans, Chamaesyce simulans
Name authority	Linnaeus: Sp. Pl. 1: 454. (1753)	(L. C. Wheeler) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Euphorbia simulans

gulf sandmat

mimicking sandmat, similar spurge

Habit

Herbs, annual or perennial, with taproot.

Herbs, annual or short-lived perennial, with usually slender, occasionally slightly thickened, rootstock.

Stems

prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose.

prostrate to reclining, 5–40 cm, glabrous.

Leaves

opposite;

stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose;

petiole 0.5–1 mm, sparsely strigose-tomentulose;

blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally.

opposite;

stipules distinct, subulate, 0.5–0.7 mm, glabrous;

petiole 0.7–1.3 mm, glabrous;

blade orbiculate, oval, to shortly oblong, 1–3.2 × 1.5–5 mm, base subsymmetric, rounded, margins entire, apex usually rounded, occasionally emarginate, surfaces glabrous;

venation obscure, only midvein conspicuous.

Involucre

broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose;

glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm;

appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate.

turbinate to campanulate, 0.8–1.2 × 0.7–1 mm, glabrous;

glands 4, red to purple, slightly concave, elliptic, 0.2–0.3 × 0.4–0.5 mm;

appendages absent.

Staminate flowers

3–5.

15–36.

Pistillate flowers

ovary densely strigose;

styles 0.4–0.5 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.2–0.3 mm, 2-fid nearly entire length.

Capsules

conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose;

columella 0.6–1 mm.

broadly ovoid, 1.3–1.8 × 1.5–2 mm, glabrous;

columella 1.1–1.4 mm.

Seeds

white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate.

whitish, reddish brown beneath coat, oblong, 4-angled in cross section, 1.5–2 × 1.3–1.8 mm, with 5–7 faint transverse ridges or wrinkles.

Cyathia

usually in small, cymose clusters on congested, axillary branches;

peduncle (0–)0.1–0.3 mm.

solitary at distal nodes;

peduncle 0.3–0.7 mm.

= 28.

Euphorbia thymifolia

Euphorbia simulans

Phenology

Flowering and fruiting midsummer–early fall.

Flowering and fruiting year-round.

Habitat

Disturbed areas, often near salt water.

Desert scrub, mountains, hills, canyons, arroyos, flats, roadsides, clay, sandy, gravelly, and rocky soils.

Elevation

0–20 m. (0–100 ft.)

600–1300 m. (2000–4300 ft.)

Distribution

FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia]

[BONAP county map]

TX; Mexico (Chihuahua, Coahuila)

[BONAP county map]

Discussion

Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia simulans, which in the flora area is known only from Brewster, Hudspeth, and Presidio counties, is difficult to distinguish in the field from the sympatric E. theriaca var. theriaca, because they are mainly distinguished by seed morphology. The latter has smaller seeds with (two or) three (or four) prominent transverse ridges, whereas E. simulans has larger seeds that are slightly wrinkled.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 12, p. 291.

FNA vol. 12, p. 289.

Parent taxa

Euphorbiaceae > Euphorbia > sect. Anisophyllum

Sibling taxa

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

Synonyms

Chamaesyce thymifolia

E. polycarpa var. simulans, Chamaesyce simulans

Name authority

Linnaeus: Sp. Pl. 1: 454. (1753)

(L. C. Wheeler) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Euphorbia thymifolia

Euphorbia simulans

Euphorbia thymifolia

Euphorbia simulans