Euphorbia thymifolia |
Euphorbia perennans |
|
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gulf sandmat |
perennial sandmat, Terlingua spurge |
|
Habit | Herbs, annual or perennial, with taproot. | Herbs, perennial, with strongly thickened, woody rootstock. |
Stems | prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose. |
erect, 7–45 cm, glabrous. |
Leaves | opposite; stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose; petiole 0.5–1 mm, sparsely strigose-tomentulose; blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally. |
opposite; stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous; petiole 0.8–2 mm, glabrous; blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous. |
Involucre | broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose; glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate. |
broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous; glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm; appendages absent. |
Staminate flowers | 3–5. |
35–45. |
Pistillate flowers | ovary densely strigose; styles 0.4–0.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.7–0.9 mm, 2-fid nearly entire length. |
Capsules | conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose; columella 0.6–1 mm. |
subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous; columella 2.2–2.7 mm. |
Seeds | white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate. |
white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled. |
Cyathia | usually in small, cymose clusters on congested, axillary branches; peduncle (0–)0.1–0.3 mm. |
solitary at distal nodes or at nodes of short, axillary branches; peduncle 1.8–3 mm. |
Euphorbia thymifolia |
Euphorbia perennans |
|
Phenology | Flowering and fruiting midsummer–early fall. | Flowering and fruiting spring–fall. |
Habitat | Disturbed areas, often near salt water. | Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. |
Elevation | 0–20 m. (0–100 ft.) | 900–1200 m. (3000–3900 ft.) |
Distribution |
FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia] |
TX; Mexico (Chihuahua) |
Discussion | Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011). Euphorbia perennans is known in the flora area only from Brewster County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 291. | FNA vol. 12, p. 282. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce thymifolia | Chamaesyce perennans |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) |
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