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gulf sandmat

Laredo sandmat

Habit Herbs, annual or perennial, with taproot. Herbs, annual, with taproot.
Stems

prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose.

prostrate, ± mat-forming, 10–20 cm, densely ashy pilose-tomentose.

Leaves

opposite;

stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose;

petiole 0.5–1 mm, sparsely strigose-tomentulose;

blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally.

opposite;

stipules distinct, filiform, 0.5–1 mm, pilose-tomentose;

petiole 0.5–1 mm, pilose-tomentose;

blade ovate to elliptic-oblong, 3–6 × 3–5 mm, base markedly asymmetric, rounded to slightly auriculate, margins usually entire, rarely largest leaves sparsely serrulate, apex acute to obtuse, surfaces moderately to densely strigose; 3-veined from base.

Involucre

broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose;

glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm;

appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate.

obconic, 0.6–1 × 0.5–1 mm, densely strigose;

glands 4, yellowish to reddish, oval to oblong, 0.1 × 0.2–0.3 mm;

appendages white to pink, rudimentary or minute, (0–)0.1–0.2 × (0–)0.1–0.3 mm, distal margin crenulate.

Staminate flowers

3–5.

3–5.

Pistillate flowers

ovary densely strigose;

styles 0.4–0.5 mm, 2-fid 1/2 length.

ovary densely white villous;

styles 0.1–0.2 mm, 2-fid 1/2 length.

Capsules

conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose;

columella 0.6–1 mm.

broadly ovoid, 1.3–1.5 × 1.4–1.5 mm, villous on keels, often glabrous or less hairy between keels;

columella 1.1–1.3 mm.

Seeds

white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate.

white, barely concealing brown undercoat, 4-angled, sharply angled in cross section, abaxial faces plane to convex, adaxial faces concave, 1.1–1.2 × 0.5–0.7 mm, with several rounded, irregular, transverse ridges.

Cyathia

usually in small, cymose clusters on congested, axillary branches;

peduncle (0–)0.1–0.3 mm.

solitary or in small, cymose clusters at distal nodes or on congested, axillary branches;

peduncle 0.5–1.5 mm.

Euphorbia thymifolia

Euphorbia laredana

Phenology Flowering and fruiting midsummer–early fall. Flowering and fruiting almost year-round.
Habitat Disturbed areas, often near salt water. Open sandy, loamy, or gravelly sites, old dunes, pastures.
Elevation 0–20 m. (0–100 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia]
[BONAP county map]
from FNA
TX; Mexico (Tamaulipas)
[BONAP county map]
Discussion

Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia laredana is similar to E. prostrata but differs from that species in its more densely tomentose indumentum, leaves with usually entire rather than serrulate margins, and slightly longer seeds with rounded rather than sharp ridges. The species occurs primarily in southern Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 291. FNA vol. 12, p. 274.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce thymifolia Chamaesyce laredana
Name authority Linnaeus: Sp. Pl. 1: 454. (1753) Millspaugh: Pittonia 2: 88. (1890)
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